Animantarx
Reconstructed skeleton
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Clade: Dinosauria
Clade: †Ornithischia
Clade: †Thyreophora
Suborder: †Ankylosauria
Family: †Nodosauridae
Subfamily: †Nodosaurinae
Clade: †Panoplosaurini
Genus: †Animantarx
Carpenter et al. 1999
Type species
†Animantarx ramaljonesi

Animantarx (/ænɪˈmæntɑːrks/ ann-i-MAN-tarks; meaning 'living citadel') is a genus of nodosaurid ankylosaurian dinosaur from the Early and Late Cretaceous of western North America. Like other nodosaurs, it would have been a slow-moving quadrupedal herbivore covered in heavy armor scutes, but without a tail club. The skull measures approximately 25 cm (10 inches) in length, suggesting the animal as a whole was no more than 3 meters (10 feet) long.

Discovery and species

Detail of skull and neck. Reconstructed skeleton at the USU Eastern Prehistoric Museum, Price, Utah

The generic name is composed of the Latin words animatus ("living" or "animated") and arx ("fortress" or "citadel"), referring to its armored nature. In particular, the name is a reference to a comment made by paleontologist R. S. Lull about ankylosaurs, that as "an animated citadel, these animals must have been practically unassailable..."[1] The type species is the only one known so far, and is called A. ramaljonesi after its discoverer, Ramal Jones. His wife, Carol Jones, also discovered the contemporaneous dinosaur Eolambia nearby.

Only one specimen of Animantarx has so far been recovered. The remains include the lower jaw and back half of the skull, along with neck and back vertebrae, and various limb elements. Animantarx is characterized by a unique combination of features, including a highly domed skull back, small horns on the postorbital and quadratojugal bones of the skull, and a mandible which is only armoured on half of its length.

Fossils in this region are often slightly radioactive, and remains of Animantarx were actually discovered following a radiological survey of the area performed by Ramal Jones, which located a higher level of radioactivity at a certain location. Subsequent excavation at this site turned up the fossil skeleton of Animantarx; no bones had been exposed on the surface.[2]

Classification

Size comparison with life restoration

Animantarx is universally thought of as a nodosaurid ankylosaur, although its precise relationships within that family are uncertain. The most recent cladistic analysis of ankylosaur phylogeny does not include Animantarx, although the authors recognize the genus as Nodosauridae incertae sedis because of its rounded supraorbital protrusions and a "knoblike" acromion on the scapula.[3] Two separate studies have found Animantarx to be the sister taxon of Edmontonia within Nodosauridae.[4][5]

The Cladgram below follows the 2018 phylogenetic analysis of Rivera-Sylva and colleagues, limited to the relationships within Panoplosaurini.[6][7]

Panoplosaurini

Animantarx

Panoplosaurus

Patagopelta

Texasetes

Denversaurus

Edmontonia longiceps

Edmontonia rugosidens

Paleoenvironment

Other angle of skeleton

These fossil remains were discovered in the Mussentuchit Member of the Cedar Mountain Formation in the eastern portion of the U.S. state of Utah. This section of the formation is believed to represent the late Albian through early Cenomanian stages of the Late Cretaceous Period, or about 106 to 97 million years ago. At least 80 other vertebrate species are known from the Mussentuchit, including fish, frogs, lizards, snakes, crocodilians, dinosaurs, birds, and mammals, although not all are complete enough to name. Many dinosaur groups are represented by fossils from this member, including carnivorous theropods as well as several different herbivorous types, including the iguanodont Eolambia. The presence of aquatic animals like fish and frogs, as well as the mudstone in which their fossils are found, suggests that this was a floodplain environment.[8]

Earlier layers within the Cedar Mountain Formation contain different nodosaur species. The oldest layer, known as the Yellow Cat Member, contains Gastonia burgei, the intermediate Poison Strip Member contains remains which may belong to Sauropelta, and the younger Ruby Ranch Member contains remains of a second species of Gastonia, G. lorriemcwhinneyae.[9] The Mussentuchit Member, which is the youngest member of the Cedar Mountain, contains Animantarx and Peloroplites.[10] While there is still a lot of exploration left to be done, this division of nodosaur species corresponds with that of other dinosaur groups and provides support for the hypothesis of three separate faunas in the Cedar Mountain Formation. The Mussentuchit fauna includes many taxa which may be of Asian origin and suggests a dispersal event may have occurred from Asia into North America around this time.[8][11]

See also

References

  1. ↑ Lull, R.S. 1914. Rulers of the Mesozoic. Yale Review 3: 352-363.
  2. ↑ Jones, R.D. & Burge, D.L. 1995. Radiological surveying as a method for mapping dinosaur bone sites. Journal of Vertebrate Paleontology 15: 38A.
  3. ↑ Vickaryous, M.K., Maryanska, T., & Weishampel, D.B. 2004. Ankylosauria. In: Weishampel, D.B., Dodson, P., & Osmólska, H. (Eds.). The Dinosauria (2nd edition). Berkeley: University of California Press. Pp. 363-392.
  4. ↑ Carpenter, K. 2001. Phylogenetic analysis of the Ankylosauria. In: Carpenter, K. (Ed.). The Armored Dinosaurs. Bloomington: Indiana University Press. Pp. 454–483.
  5. ↑ Hill, R.V., Witmer, L.M., Norell, M.A. 2003. A New specimen of Pinacosaurus grangeri (Dinosauria: Ornithischia) from the Late Cretaceous of Mongolia: ontogeny and phylogeny of ankylosaurs. American Museum Novitates 3395: 1-29.
  6. ↑ Rivera-Sylva, Héctor E.; Frey, Eberhard; Stinnesbeck, Wolfgang; Carbot-Chanona, Gerardo; Sanchez-Uribe, Iván E.; Guzmán-Gutiérrez, José Rubén (2018). "Paleodiversity of Late Cretaceous Ankylosauria from Mexico and their phylogenetic significance". Swiss Journal of Palaeontology. 137 (1): 83–93. doi:10.1007/s13358-018-0153-1. ISSN 1664-2384. S2CID 134924657.
  7. ↑ Madzia, D.; Arbour, V.M.; Boyd, C.A.; Farke, A.A.; Cruzado-Caballero, P.; Evans, D.C. (2021). "The phylogenetic nomenclature of ornithischian dinosaurs". PeerJ. 9: e12362. doi:10.7717/peerj.12362. PMC 8667728. PMID 34966571.
  8. 1 2 Kirkland, J.I., Britt, B., Burge, D.L., Carpenter, K., Cifelli, R., DeCourten, F., Eaton, J., Hasiotis, S., and Lawton, T. 1997. Lower to Middle Cretaceous dinosaur faunas of the Central Colorado Plateau: a key to understanding 35 million years of tectonics, sedimentology, evolution, and biogeography. Brigham Young University Geology Studies 42:69-103.
  9. ↑ Kinneer, B.; Carpenter, K.; Shaw, A. (2016). "Redescription of Gastonia burgei (Dinosauria: Ankylosauria, Polacanthidae), and description of a new species". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 282 (1): 37–80. doi:10.1127/njgpa/2016/0605.
  10. ↑ Carpenter, Kenneth; Bartlett, Jeff; Bird, John; Barrick, Reese (2008). "Ankylosaurs from the Price River Quarries, Cedar Mountain Formation (Lower Cretaceous), east-central Utah". Journal of Vertebrate Paleontology. 28 (4): 1089–1101. doi:10.1671/0272-4634-28.4.1089. S2CID 129480044.
  11. ↑ Carpenter, K., Kirkland, J.I., Burge, D.L., & Bird, J. 1999. Ankylosaurs (Dinosauria: Ornithischia) of the Cedar Mountain Formation, Utah, and their stratigraphic distribution. In: Gillette, D. (Ed.) Vertebrate Paleontology in Utah. Utah Geological Survey Miscellaneous Publication 99-1. Pp. 243-251.
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