| Icerya | |
|---|---|
![]()  | |
| Icerya purchasi, female | |
| Scientific classification  | |
| Domain: | Eukaryota | 
| Kingdom: | Animalia | 
| Phylum: | Arthropoda | 
| Class: | Insecta | 
| Order: | Hemiptera | 
| Suborder: | Sternorrhyncha | 
| Family: | Monophlebidae | 
| Genus: | Icerya Signoret, 1875  | 
| Species | |
| 
 See text  | |
Icerya is a genus of scale insects in the family Monophlebidae. It is named after physician-naturalist Dr. Edmond Icery of British Mauritius.[1]
Hermaphroditism
Hermaphroditism is extremely rare in the insect world despite the comparatively common nature of this condition in the crustaceans. Several species of Icerya, including the pestiferous cottony-cushion scale, I. purchasi, are known to be hermaphrodites that reproduce by self-fertilising. Occasionally males are produced from unfertilised eggs, but generally individuals are monoecious with a female-like nature but possessing an ovotestis (a part-testis, part-ovary organ) and sperm is transmitted ovarially from the female to her young.[2] The existence of both hermaphrodites and males in a species is known as androdioecy. This hermaphroditic sexual self-sufficiency, where a single individual can populate new territory, has contributed to the invasive spread of the cottony-cushion scale insect away from its native Australia.[3]
List of species
- Icerya aegyptiaca (Douglas 1890).
 - Icerya albolutea Cockerell 1898.
 - Icerya bimaculata De Lotto 1959.
 - Icerya brachystegiae Hall 1940.
 - Icerya brasiliensis Hempel 1920.
 - Icerya callitri (Froggatt 1921).
 - Icerya chilensis Hempel 1920.
 - Icerya colimensis Cockerell 1902.
 - Icerya flava Hempel 1920.
 - Icerya flocculosa Hempel 1932.
 - Icerya formicarum Newstead 1897.
 - Icerya genistae Hempel 1912.
 - Icerya hanoiensis Jashenko & Danzig 1992.
 - Icerya imperatae Rao 1951.
 - Icerya insulans Hempel 1923.
 - Icerya koebelei Maskell 1892.
 - Icerya leuderwaldti Hempel 1918.
 - Icerya littoralis mimosae Cockerell 1902.
 - Icerya littoralis tonilensis Cockerell 1902.
 - Icerya littoralis Cockerell 1898.
 - Icerya longisetosa Newstead 1911.
 - Icerya maxima Newstead 1915.
 - Icerya maynei Vayssiere 1926.
 - Icerya menoni Rao 1951.
 - Icerya minima Morrison 1919.
 - Icerya minor Green 1908.
 - Icerya montserratensis Riley & Howard 1890.
 - Icerya morrisoni Rao 1951.
 - Icerya nigroareolata Newstead 1917.
 - Icerya palmeri Riley & Howard 1890.
 - Icerya paulista Hempel 1920.
 - Icerya pilosa Green 1896.
 - Icerya pulchra (Leonardi 1907).
 - Icerya purchasi citriperda Hempel 1920.
 - Icerya purchasi crawii Cockerell 1897.
 - Icerya purchasi maskelli Cockerell 1897.
 - Icerya purchasi Maskell 1878 - cottony cushion scale
 - Icerya rileyi Cockerell 1896.
 - Icerya schoutedeni Vayssiere 1926.
 - Icerya schrottkyi Hempel 1900.
 - Icerya seychellarum (type) (Westwood 1855).
 - Icerya seychellarum cristata Newstead 1909.
 - Icerya similis Morrison 1923.
 - Icerya splendida Lindinger 1913.
 - Icerya subandina Leonardi 1911.
 - Icerya sulfurea pattersoni Newstead 1917.
 - Icerya sulfurea Lindinger 1913.
 - Icerya sumatrana Rao 1951.
 - Icerya taunayi Hempel 1920.
 - Icerya travancorensis Rao 1951.
 - Icerya tremae Vayssiere 1926.
 - Icerya zeteki Cockerell 1914.
 - Icerya zimmermani Green 1932.
 
References
- ↑ Sorensen, W. Conner; Smith, Edward H. (2019). "Vedalia the "Wonder Beetle" and Biological Control". Charles Valentine Riley: Founder of Modern Entomology. And Janet R. Smith, with Donald C. Weber. Tuscaloosa: University of Alabama Press. p. 221. ISBN 9780817392222.
 - ↑ Normark, Benjamin B. (2003). "The Evolution of Alternative Genetic Systems in Insects". Annual Review of Entomology. 48 (1): 397–423. doi:10.1146/annurev.ento.48.091801.112703. ISSN 0066-4170.
 - ↑ The Insects An outline of Entomology, Gullan & Cranston, Wiley-Blackwell 2001
 
