List of years in paleobotany
In paleontology
2016
2017
2018
2019
2020
2021
2022
In arthropod paleontology
2016
2017
2018
2019
2020
2021
2022
In paleoentomology
2016
2017
2018
2019
2020
2021
2022
In paleomalacology
2016
2017
2018
2019
2020
2021
2022
In reptile paleontology
2016
2017
2018
2019
2020
2021
2022
In archosaur paleontology
2016
2017
2018
2019
2020
2021
2022
In mammal paleontology
2016
2017
2018
2019
2020
2021
2022
In paleoichthyology
2016
2017
2018
2019
2020
2021
2022

This article records new taxa of fossil plants that are scheduled to be described during the year 2019, as well as other significant discoveries and events related to paleobotany that are scheduled to occur in the year 2019.

Mosses

Name Novelty Status Authors Age Type locality Location Notes Images

Heinrichsiella[1]

Gen. et sp. nov

Valid

Bippus et al.

Jurassic

 Argentina

A moss, possibly related to the family Polytrichaceae or Timmiellaceae. Genus includes new species H. patagonica.

Kulindobryum[2]

Gen. et sp. nov

Valid

Ignatov in Mamontov & Ignatov

Middle or Late Jurassic

Ukureyskaya Formation

 Russia
( Zabaykalsky Krai)

A form genus of dispersed moss capsules. Genus includes new species K. taylorioides.

Polycingulatisporites multiverrucata[3]

Sp. nov

In press

Santamarina in Santamarina et al.

Late Cretaceous (Cenomanian)

Mata Amarilla Formation

 Argentina

Spores of a member of Bryophyta of uncertain phylogenetic placement, possibly of sphagnaceous affinity. Announced in 2019; the final version was scheduled to be published in 2020.

Sphagnum heinrichsii[4]

Sp. nov

Valid

Ignatov et al.

Late Eocene

Rovno amber

 Ukraine

A moss, a species of Sphagnum.

Paleaethallus[2]

Gen. et sp. nov

Valid

Mamontov, Katagiri & Borovich in Mamontov & Ignatov

Late Jurassic

Glushkovo Formation

 Russia
( Zabaykalsky Krai)

A thalloid bryophyte. Genus includes new species P. squarrosus.

Liverworts

Name Novelty Status Authors Age Type locality Location Notes Images

Khasurtythallus[2]

Gen. et sp. nov

Valid

Mamontov in Mamontov & Ignatov

Early Cretaceous

 Russia
( Buryatia)

A Marchantiidae liverwort.
The type species is K. monosolenioides.

Ricciopsis sandaolingensis[5]

Sp. nov

Valid

Li & Sun in Li et al.

Middle Jurassic

Xishanyao Formation

 China

A Ricciaceae liverwort.

Thallites yangcaogouensis[6]

Sp. nov

Valid

Wang et al.

Late Triassic

Yangcaogou Formation

 China

A plant of uncertain phylogenetic placement, probably a liverwort.

Ferns and fern allies

Name Novelty Status Authors Age Type locality Location Notes Images

Alloiopteris loecsei[7]

Sp. nov

Valid

Pšenička et al.

Carboniferous (Moscovian)

 Germany

A zygopterid fern.

Annularia noronhai[8]

Sp. nov

Valid

Correia et al.

Carboniferous (Gzhelian)

Douro Basin

 Portugal

A member of the family Calamitaceae. Announced in 2019; the final version of the article naming it was published in 2021.

Azolla keuja[9]

Sp. nov

Valid

Jud, De Benedetti, Gandolfo & Hermsen

Paleocene (Danian)

Salamanca Formation

 Argentina

A species of Azolla.

Berendtiopteris[10]

Gen. et comb. nov

Valid

Sadowski et al.

Eocene

Baltic amber

Europe (Baltic Sea region)

A plant of uncertain phylogenetic placement, probably a fern; a new genus for "Pecopteris" humboldtiana.

Bifariusotheca[11]

Gen. et sp. nov

Valid

X.H.Zhao ex Doweld

Late Permian

Longtan Formation

 China

A marattialean fern. Genus includes new species B. notocathaysica Doweld.

Bowmanites yongchangensis[12]

Sp. nov

Valid

Sun et al.

Permian (Cisuralian)

 China

A member of Sphenophyllales.

Clavatisporites cenomaniana[3]

Sp. nov

In press

Santamarina in Santamarina et al.

Late Cretaceous (Cenomanian)

Mata Amarilla Formation

 Argentina

Spores of a member of Filicopsida of uncertain phylogenetic placement. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.

Collarisporites minor[3]

Sp. nov

In press

Santamarina in Santamarina et al.

Late Cretaceous (Cenomanian)

Mata Amarilla Formation

 Argentina

Spores of a member of Filicopsida of uncertain phylogenetic placement. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.

Cyathocarpus yongchangensis[13]

Sp. nov

Valid

Sun & Sun in Sun et al.

Permian (Cisuralian)

Shanxi Formation

 China

A fern related to Psaronius.

Floratheca[14]

Gen. et sp. nov

Valid

Lundgren et al.

Early Permian

Río Genoa Formation

 Argentina

A member of Marattiales. Genus includes new species F. apokalyptika.

Germera brousmicheae[15]

Sp. nov

Valid

Álvarez-Vázquez

Carboniferous (Westphalian)

 Canada

A member of Filicopsida.

Hausmannia olaensis[16]

Sp. nov

Valid

Golovneva & Grabovskiy

Late Cretaceous (Santonian–early Campanian)

 Russia

A Dipteridaceae umbrella fern.

Heinrichsia[17]

Gen. et sp. nov

Valid

Regalado et al.

Cretaceous

Burmese amber

 Myanmar

A fern belonging to the family Pteridaceae. Genus includes new species H. cheilanthoides.

Kamatheca[11]

Gen. et comb. nov

Valid

Doweld

Permian

 Russia

A marattialean fern; a new genus for "Acitheca" gigantea Esaulova.

Marsilea sprungerorum[18]

Sp. nov

Valid

Hermsen

Eocene

Green River Formation

 United States
( Colorado
 Utah)

A Marsilea species water fern.

Neolobatannularia[19]

Gen. et sp. nov

Valid

Sun & Li in Wang et al.

Late Triassic

Yangcaogou Formation

 China

A member of Equisetales. Genus includes new species N. liaoningensis.

Osmundastrum gvozdevae[20]

Sp. nov

Valid

Bazhenova & Bazhenov

Middle Jurassic (Bathonian)

 Russia
( Kursk Oblast)

A species of Osmundastrum.

Palaeosorum waipiata[21]

Sp. nov

Valid

Kaulfuss et al.

Early Miocene

 New Zealand

A member of the family Polypodiaceae.

Phlebopteris kirchneri[22]

Sp. nov

Valid

Barbacka & Kustatscher in Barbacka, Kustatscher & Bodor

Early Jurassic (Hettangian)

Mecsek Coal Formation

 Hungary

A fern belonging to the family Matoniaceae.

Plenasium xiei[23]

Sp. nov

Valid

Cheng et al.

Cretaceous

 China

A member of Osmundaceae. Announced in 2019; the final version of the article naming it was published in 2021.

Polymorphopteris magdalenae[11]

Sp. nov

Valid

R.H.Wagner ex Doweld

Late Carboniferous

 Spain

A marattialean fern.

Polymorphopteris wagneri[11]

Sp. nov

Valid

Doweld

Late Carboniferous (Kasimovian)

 Spain

A marattialean fern.

Polypodiisporites serratus[24]

Sp. nov

Valid

D'Apolito et al.

Pliocene–Pleistocene

 Brazil

Fossil spore of a member of the family Polypodiaceae.

Polypodiisporites timidus[24]

Sp. nov

Valid

D'Apolito et al.

Pliocene–Pleistocene

 Brazil

Fossil spore of a member of the family Polypodiaceae.

Rinistachya[25]

Gen. et sp. nov

Valid

Prestianni & Gess

Devonian (Famennian)

Witpoort Formation

 South Africa

A member of Sphenophyllales.
The type species is R. hilleri.

Rothwellopteris[26]

Gen. et sp. nov

Valid

He et al.

Late Permian

Xuanwei Formation

 China

A fern belonging to the group Marattiales. Genus includes new species R. pecopteroides.

Scolecopteris libera[27]

Sp. nov

Valid

Li et al.

Permian (Asselian)

Taiyuan Formation

 China

A marattialean fern

Scolecopteris renaultii[11]

Sp. nov

Valid

Doweld

Permian (Cisuralian)

 France

A marattialean fern.

Tiania resinus[28]

Sp. nov

Valid

He & Wang

Permian (Lopingian)

Xuanwei Formation

 China

A member of Osmundales belonging to the extinct family Guaireaceae.

Lycophytes

Name Novelty Status Authors Age Type locality Location Notes Images

Bergeria wenquanensis[29]

Sp. nov

Valid

Feng, D’Rozario & Zhang

Carboniferous (Viséan)

Akeshake Formation

 China

A member of Lepidodendrales belonging to the family Flemingitaceae.

Guangdedendron[30]

Gen. et sp. nov

Wang et al.

Devonian (Famennian)

Wutong Formation

 China

A member of Isoetales belonging to the group Dichostrobiles. Genus includes new species G. micrum.

Omphalophloios wagneri[31]

Sp. nov

Valid

Opluštil, Pšenička & Bek

Carboniferous (Moscovian)

Illinois Basin

 United States
( Indiana)

Sawdonia hippotheca[32]

Sp. nov

Valid

Berry & Gensel

Devonian (probably late Givetian)

Campo Chico Formation

 Venezuela

A member of Zosterophyllopsida.

Conifers

Araucariaceae

Name Novelty Status Authors Age Type locality Location Notes Images

Agathoxylon gilii[33]

Sp. nov

Valid

Ríos-Santos & Cevallos-Ferriz

Late Jurassic

Todos Santos Formation

 Mexico

Agathoxylon hoodii[34]

Comb nov

valid

(Tidwell & Medlyn) Gee et al

Late Jurassic

Morrison Formation

 USA
 Utah

An araucariaceous petrified wood.
Moved from Araucarioxylon hoodii (1993)[35]

Agathoxylon jericonse[33]

Sp. nov

Valid

Ríos-Santos & Cevallos-Ferriz

Late Jurassic

Todos Santos Formation

 Mexico

Agathoxylon kotaense[36]

Sp. nov

In press

Chinnappa, Rajanikanth & Pauline Sabina

?Late JurassicEarly Cretaceous

Kota Formation

 India

A member of the family Araucariaceae.

Agathoxylon parrensis[33]

Sp. nov

Valid

Ríos-Santos & Cevallos-Ferriz

Paleocene

Las Encinas Formation

 Mexico

Araucaria balfourensis[37]

Sp. nov

Valid

Hill et al.

Cenozoic

 Australia

A species of Araucaria.

Araucaria macrophylla[37]

Sp. nov

Valid

Hill et al.

Cenozoic

 Australia

A species of Araucaria.

Araucaria mollifolia[37]

Sp. nov

Valid

Hill et al.

Cenozoic

 Australia

A species of Araucaria.

Araucaria rothwellii[38]

Sp. nov

Valid

Kvaček in Kvaček et al.

Late Cretaceous (Campanian-Maastrichtian)

Bozova Formation

 Turkey

A species of Araucaria.

Brachyphyllum garciarum[39]

Sp. nov

Valid

Carrizo et al.

Early Cretaceous (early Hauterivian/early Barremian)

Springhill Formation

 Argentina

Probably a member of the family Araucariaceae.

Cupressaceae

Name Novelty Status Authors Age Type locality Location Notes Images

Austrocupressinoxylon[40]

Gen. et sp. nov

Valid

Nunes et al.

Early Cretaceous

 Argentina

A member of Cupressaceae. Genus includes new species A. barcinense.

Austrohamia asfaltensis[41]

Sp. nov

Valid

Contreras et al.

Early Jurassic

Cañadón Asfalto Formation

 Argentina

A member of the family Cupressaceae.

Callitris blackburnii[42]

Sp. nov

Valid

Paull et al.

Middle Miocene

 Australia

A species of Callitris.

Cupressinoxylon pliocenica[43]

Sp. nov

Valid

Akkemik

Pliocene

Örencik Formation

 Turkey

A member of the family Cupressaceae described on the basis of fossil wood.

Mesocyparis sinica[44]

Sp. nov

Valid

Cui et al.

Paleocene (Danian)

Wuyun Formation

 China

A member of the family Cupressaceae.

Protaxodioxylon sahnii[45]

Sp. nov

Valid

Chinnappa, Kavali & Rajanikanth

Late Jurassic to Early Cretaceous

Kota Formation

 India

A member of Cupressaceae, possibly related to Taxodium.

Protodammara reimatamoriori[46]

Sp. nov

Valid

Mays & Cantrill

Late Cretaceous (Cenomanian)

Tupuangi Formation

 New Zealand

A member of Cupressaceae.

Taxodioxylon cabullensis[33]

Sp. nov

Valid

Ríos-Santos & Cevallos-Ferriz

Late Cretaceous

Packard Formation

 Mexico

A cupressaceous fossil wood.

Taxodium viligense[47]

Sp. nov

Valid

Golovneva

Late Cretaceous (Coniacian)

Chingandzha Formation

 Russia

A species of Taxodium.

Pinceae

Name Novelty Status Authors Age Type locality Location Notes Images

Abies cuitlahuacii[48]

Sp. nov

Valid

Cevallos-Ferriz, Ríos-Santos & Lozano-García

Pleistocene

 Mexico

A fir.

Pinus plioarmandii[49]

Sp. nov

Valid

An et al.

Pliocene

 China

A pine.

Pinuxylon alonissianum[50]

Sp. nov

Valid

Mantzouka & Sakala in Mantzouka et al.

Early Miocene

 Greece

A member of the family Pinaceae described on the basis of fossil wood.

Schizolepidopsis borealis[51]

Sp. nov

Valid

Domogatskaya & Herman

Early Cretaceous (Albian)

Balyktakh Formation

 Russia

A member of the family Pinaceae.

Podocarpaceae

Name Novelty Status Authors Age Type locality Location Notes Images

Dacrycarpus guipingensis[52]

Sp. nov

Valid

Wu et al.

Miocene

Erzitang Formation

 China

A species of Dacrycarpus. Announced in 2019; the final version of the article naming it was published in 2021.

Kirketapel[53]

Gen. et sp. nov

Valid

Andruchow-Colombo et al.

Paleocene (early Danian)

Salamanca Formation

 Argentina

A member of the family Podocarpaceae.
The type species is K. salamanquensis.

Podocarpus pliomacrophyllus[54]

Sp. nov

In press

Chen et al.

Early Pliocene

 China

A species of Podocarpus. Announced in 2019; the final version of the article naming it is not published yet.

Other conifers

Name Novelty Status Authors Age Type locality Location Notes Images

Cephalotaxus maguanensis[55]

Sp. nov

Valid

Zhang et al.

Middle Miocene

 China

A species of Cephalotaxus.

Cupressinocladus shelikhovii[47]

Sp. nov

Valid

Golovneva

Late Cretaceous (Coniacian)

Chingandzha Formation

 Russia

A cheirolepidiaceous species

Frenelopsis justae[56]

Sp. nov

Valid

Barral et al.

Early Cretaceous (Albian)

Escucha Formation

 Spain

A member of the family Cheirolepidiaceae.

Ningxiaites shitanjingensis[57]

Sp. nov

Valid

Wei et al.

Permian (Changhsingian)

Sunjiagou Formation

 China

A conifer wood.

Protocupressinoxylon carrizalense[58]

Sp. nov

Valid

Correa et al.

Late Triassic

Carrizal Formation

 Argentina

Other seed plants

Name Novelty Status Authors Age Type locality Location Notes Images

Amyelon bogdense[59]

Sp. nov

Valid

Wan, Yang & Wang

Late Permian or Early Triassic

Guodikeng Formation

 China

A silicified gymnospermous root.

Arazedispermum[60]

Gen. et sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous (late Aptian-early Albian)

Figueira da Foz Formation

 Portugal

A seed plant belonging to the informal grouping Bennettitales-Erdtmanithecales-Gnetales. Genus includes new species A. lustanicum.

Axsmithia[61]

Gen. et comb. nov

Valid

Anderson et al.

Triassic

Antarctica

A seed fern. Genus includes "Umkomasia" uniramia Axsmith et al. (2000).

Bowenia johnsonii[62]

Sp. nov

Valid

Hill et al.

Early Eocene

 Australia

A cycad, a species of Bowenia.

Brinkia[63]

Gen. et 2 sp. nov

Valid

Kustatscher, Visscher & van Konijnenburg-van Cittert

Permian (Lopingian)

Bellerophon Formation
Gröden/Val Gardena Sandstone

 Italy

A possible member of Czekanowskiales. Genus includes new species B. kerpiana and B. cortianensis.

Cordaabaxicutis jacobii[64]

Sp. nov

Valid

Šimůnek

Carboniferous (Pennsylvanian)

 Czech Republic

A member of Cordaitales.

Cordaadaxicutis detmarovicensis[64]

Sp. nov

Valid

Šimůnek

Carboniferous (Pennsylvanian)

 Czech Republic

A member of Cordaitales.

Cordaadaxicutis doubravensis[64]

Sp. nov

Valid

Šimůnek

Carboniferous (Pennsylvanian)

 Czech Republic

A member of Cordaitales.

Cordaadaxicutis jaroslavii[64]

Sp. nov

Valid

Šimůnek

Carboniferous (Pennsylvanian)

 Czech Republic

A member of Cordaitales.

Cordaadaxicutis orlovensis[64]

Sp. nov

Valid

Šimůnek

Carboniferous (Pennsylvanian)

 Czech Republic

A member of Cordaitales.

Cryptokerpia[65]

Gen. et sp. nov

Valid

Blomenkemper, Abu Hamad & Bomfleur

Late Permian

Umm Irna Formation

 Jordan

An enigmatic type of gymnosperm leaf. Genus includes new species C. sarlaccophora.

Douropteris[66]

Gen. et sp. nov

Valid

Correia et al.

Carboniferous (Gzhelian)

Douro Basin

 Portugal

A seed fern belonging to the group Medullosales. Genus includes new species D. alvarezii.

Ephedrispermum tenuicostatum[60]

Sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous (Aptian or early Albian)

 Portugal

A seed plant belonging to the informal grouping Bennettitales-Erdtmanithecales-Gnetales.

Geminispermum[67]

Gen. et sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous (Albian)

Potomac Group

 United States
( Virginia)

A seed plant belonging to the informal grouping Caytoniales-Umkomasiales-Petriellales. Genus includes new species G. virginiense.

Glossopteris thirroulensis[68]

Sp. nov

Valid

McLoughlin & Mays in McLoughlin, Maksimenko & Mays

Permian (Wuchiapingian)

Wilton Formation

 Australia

Hirsutisperma[69]

Gen. et sp. nov

Valid

Scott et al.

Carboniferous (Viséan)

 United Kingdom

An ovule adapted for wind dispersal and for deterring herbivory. Genus includes new species H. rothwellii.

Huncocladus[70]

Gen. et sp. nov

Valid

Andruchow-Colombo, Wilf & Escapa

Early Eocene

La Huitrera Formation

 Argentina

A seed plant of uncertain phylogenetic placement. Originally described as a member of the family Podocarpaceae related to the genus Phyllocladus; on the other hand, Dörken et al. (2021) rejected the podocarpaceous affinity of Huncocladus, and considered it to be more closely related to the cycad genera Bowenia or Eobowenia.[71][72] Genus includes new species H. laubenfelsii.

Illawarraspermum[68]

Gen. et sp. nov

Valid

McLoughlin & Mays in McLoughlin, Maksimenko & Mays

Permian (Wuchiapingian)

Wilton Formation

 Australia

A glossopterid seed. Genus includes new species I. ovatum.

Kirchmuellia[61]

Gen. et comb. nov

Valid

Anderson et al.

Early Jurassic

 Germany

A seed fern. Genus includes "Umkomasia" franconica Kirchner & Müller (1992).

Lepidopteris scassoi[73]

Sp. nov

Valid

Elgorriaga, Escapa & Cúneo

Early Jurassic

Cañadón Asfalto Formation

 Argentina

Lignieriopsis[60]

Gen. et 2 sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous (late Aptian-Albian)

Figueira da Foz Formation
Potomac Group

 Portugal
 United States
( Virginia)

A seed plant belonging to the informal grouping Bennettitales-Erdtmanithecales-Gnetales. Genus includes new species L. stenosperma and L. parva.

Mariopteris hexiensis[74]

Sp. nov

Valid

Wang et al.

Permian (Cisuralian)

Shanxi Formation

 China

Announced in 2019; the final version of the article naming it was published in 2021.

Mariopteris yongchangensis[74]

Sp. nov

Valid

Wang et al.

Permian (Cisuralian)

Shanxi Formation

 China

Announced in 2019; the final version of the article naming it was published in 2021.

Muelkirchium[75]

Gen. et comb. nov

Valid

Anderson et al.

Early Jurassic

 Germany

A seed fern. Genus includes "Pteruchus" septentrionalis Kirchner & Müller (1992).

Mutoviaspermum[76]

Gen. et sp. nov

Valid

Karasev et al.

Permian (Lopingian)

Poldarsa Formation

 Russia
( Vologda Oblast)

A member of Voltziales. Genus includes new species M. krassilovii.

Noeggerathiopsis brasiliensis[77]

Nom. nov

Valid

Degani-Schmidt & Guerra-Sommer

Early Permian

Rio Bonito Formation

 Brazil

A member of Cordaitales; a replacement name for Rufloria gondwanensis Guerra-Sommer (1989).

Potoniea krisiae[78]

Sp. nov

Valid

Pšenička, Zodrow & Bek

Carboniferous (Moscovian)

Sydney Coalfield

 Canada
( Nova Scotia)

Reproductive male organ of a seed fern, possibly a member of the family Parispermaceae.

Protophyllocladoxylon zhaobishanensis[79]

Sp. nov

Valid

Wan, Yang & Wang

Early Triassic (Induan)

Jiucaiyuan Formation

 China

A silicified gymnospermous fossil wood.

Pseudotorellia yimaensis[80]

Sp. nov

Valid

Dong et al.

Middle Jurassic

Yima Formation

 China

Ptilophyllum eminelidarum[81]

Sp. nov

Valid

Carrizo, Lafuente Diaz & Del Fueyo

Early Cretaceous

Springhill Formation

 Argentina

A member of Bennettitales.

Ptilophyllum micropapillosum[82]

Sp. nov

Valid

Lafuente Diaz et al.

Early Cretaceous

Springhill Formation

 Argentina

A member of Bennettitales.

Rothwellia[60]

Gen. et sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous (Albian)

Potomac Group

 United States
( Virginia)

A seed plant belonging to the informal grouping Bennettitales-Erdtmanithecales-Gnetales. Genus includes new species R. foveata.

Sagenopteris trapialensis[83]

Sp. nov

Valid

Elgorriaga, Escapa & Cúneo

Early Jurassic

Lonco Trapial Formation

 Argentina

A member of Caytoniales.

Sclerospiroxylon xinjiangensis[84]

Sp. nov

Valid

Wan, Yang & Wang

Permian (Kungurian)

Hongyanchi Formation

 China

Sueria laxinervis[85]

Sp. nov

Valid

Yamada & Nishida in Yamada et al.

Late Cretaceous (Maastrichtian)

Quiriquina Formation

 Chile

A cycad.

Thodaya[60]

Gen. et sp. nov

Junior homonym

Friis, Crane & Pedersen

Early Cretaceous (Albian)

Potomac Group

 United States
( Virginia)

A seed plant belonging to the informal grouping Bennettitales-Erdtmanithecales-Gnetales. Genus includes new species T. sykesiae. The generic name is preoccupied by Thodaya Compton.

Umaltolepis yimaensis[80]

Sp. nov

Valid

Dong et al.

Middle Jurassic

Yima Formation

 China

Umkomasia corniculata[86]

Sp. nov

Valid

Shi et al.

Early Cretaceous (AptianAlbian)

 Mongolia

Umkomasia trilobata[86]

Sp. nov

Valid

Shi et al.

Early Cretaceous (AptianAlbian)

 Mongolia

Wangjunia[87]

Gen. et sp. nov

Valid

Backer, Bomfleur & Kerp

Permian (Guadalupian)

Lower Shihhotse Formation

 China

A member of Cordaitales. Genus includes new species W. microphylla.

Xuanweioxylon damogouense[88]

Sp. nov

Valid

Yang et al.

Permian (Lopingian)

Xuanwei Formation

 China

A conifer stem.

Zhangwuia[89]

Gen. et sp. nov

Valid

Liu, Hou & Wang

Middle Jurassic (Callovian)

Jiulongshan Formation

 China

A reproductive organ of a seed plant of uncertain phylogenetic placement. Genus includes new species Z. mira.

Flowering plants

Basal angiosperms

Nymphaeales

Name Novelty Status Authors Age Type locality Location Notes Images

Nuphaea[90]

Gen. et sp. nov

Valid

Gee & Taylor

Eocene

Messel pit

 Germany

A member of Nymphaeaceae. Genus includes new species N. engelhardtii.

Other basal angiosperms

Name Novelty Status Authors Age Type locality Location Notes Images

Anaspermum[91]

Gen. et sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous (late Barremian-early Aptian)

Almargem Formation

 Portugal

A flowering plant with affinities to Austrobaileyales or Nymphaeales. Genus includes new species A. operculatum.

Gastonispermum antiquum[91]

Sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous (late Barremian-early Aptian)

Almargem Formation

 Portugal

A flowering plant with affinities to Austrobaileyales or Nymphaeales.

Monocots

Alismatales

Name Novelty Status Authors Age Type locality Location Notes Images

Natantisphyllum[92]

Gen. et sp. nov

Valid

Puebla, Vento & Prámparo

Late Cretaceous

 Argentina

A member of the family Araceae. Genus includes new species N. crenae. Announced in 2019; the final version of the article naming it was published in 2021.

Orontiophyllum ferreri[93]

Sp. nov

Valid

Sender et al.

Early Cretaceous (Albian)

 Spain

A member or a relative of the family Araceae.

Turolospadix[93]

Gen. et sp. nov

Valid

Sender et al.

Early Cretaceous (Albian)

 Spain

A member or a relative of the family Araceae. Genus includes new species T. bogneri.

Arecales

Name Novelty Status Authors Age Type locality Location Notes Images

Palmoxylon deoriensis[94]

Sp. nov

Valid

Khan, Mandal & Bera

Late Cretaceous (late Maastrichtian) – early Paleocene (Danian)

Deccan Intertrappean Beds

 India

A permineralized palm stem.

Sabalites tibetensis[95]

Sp. nov

Valid

Su & Zhou in Su et al.

Oligocene (Chattian)

Lunpola Basin

 China

A member of the family Arecaceae belonging to the subfamily Coryphoideae.

Sclerosperma protoprofizianum[96]

Sp. nov

Valid

Grímsson & Zetter in Grímsson et al.

Late Oligocene

 Ethiopia

A species of Sclerosperma.

Sclerosperma protomannii[96]

Sp. nov

Valid

Grímsson & Zetter in Grímsson et al.

Late Oligocene

 Ethiopia

A species of Sclerosperma.

Spinopalmoxylon cicatricosum[97]

Sp. nov

Valid

Winterscheid

Oligocene

Köln Formation

 Germany

A member of the family Arecaceae belonging to the tribe Calameae.

Spinopalmoxylon parvifructum[97]

Sp. nov

Valid

Winterscheid

Oligocene

Köln Formation

 Germany

A member of the family Arecaceae belonging to the tribe Calameae.

Spinizonocolpites riochiquensis[98]

Sp. nov

Valid

Vallati & De Sosa Tomas in Vallati, De Sosa Tomas & Casal

Late Cretaceous (Maastrichtian)

Lago Colhué Huapí Formation

 Argentina

A member of Arecaceae described on the basis of fossil pollen grains. Announced in 2019; the final version of the article naming it was published in 2020.

Dioscoreales

Name Novelty Status Authors Age Type locality Location Notes Images

Dioscorea eocenicus[99]

Sp. nov

Valid

Mehrotra & Shukla

Early Eocene

 India

A species of Dioscorea.

Dioscorea manchesteri[100]

Sp. nov

Valid

Kvaček

Miocene

Most Formation

 Czech Republic

A species of Dioscorea.

Poales

Name Novelty Status Authors Age Type locality Location Notes Images

Bambusiculmus makumensis[101]

Sp. nov

Valid

Srivastava et al.

Late Oligocene

 India

A bamboo.

Bambusiculmus tirapensis[101]

Sp. nov

Valid

Srivastava et al.

Late Oligocene

 India

A bamboo.

Bambusium arunachalense[101]

Sp. nov

Valid

Srivastava et al.

Late Miocene to Pliocene

 India

A bamboo.

Bambusium deomarense[101]

Sp. nov

Valid

Srivastava et al.

Late Miocene to Pliocene

 India

A bamboo.

Scirpus weichangensis[102]

Sp. nov

Valid

Liang in Lu et al.

Early Miocene

Hannuoba Formation

 China

A species of Scirpus.

Magnoliids

Laurales

Name Novelty Status Authors Age Type locality Location Notes Images

Cinnamomum raptiensis[103]

Sp. nov

Valid

Prasad et al.

Middle Miocene

Lower Churia Group

   Nepal

A species of Cinnamomum.

Laurinoxylon acalensis[104]

Sp. nov

Valid

Pérez-Lara, Estrada-Ruiz & Castañeda-Posadas

Eocene

El Bosque Formation

 Mexico

A member of Lauraceae.

Laurinoxylon thomasii[105]

Sp. nov

Valid

Akkemik in Akkemik, Akkılıç & Güngör

Early Miocene

 Turkey

Laurophyllum alseodaphnoides[106]

Sp. nov

Valid

Wang & Sun in Wang et al.

Miocene (Langhian)

Fotan Group

 China

A member of Lauraceae described on the basis of fossil leaves.

Laurophyllum fotanensis[106]

Sp. nov

Valid

Wang & Sun in Wang et al.

Miocene (Langhian)

Fotan Group

 China

A member of Lauraceae described on the basis of fossil leaves.

Laurophyllum lindaiensis[106]

Sp. nov

Valid

Wang & Sun in Wang et al.

Miocene (Langhian)

Fotan Group

 China

A member of Lauraceae described on the basis of fossil leaves.

Laurophyllum triangulatum[106]

Sp. nov

Valid

Wang & Sun in Wang et al.

Miocene (Langhian)

Fotan Group

 China

A member of Lauraceae described on the basis of fossil leaves.

Laurophyllum zhangpuensis[106]

Sp. nov

Valid

Wang & Sun in Wang et al.

Miocene (Langhian)

Fotan Group

 China

A member of Lauraceae described on the basis of fossil leaves.

Mezilaurinoxylon oleiferum[107]

Sp. nov

Valid

Ruiz, Brea & Pujana in Ruiz et al.

Paleocene (Danian)

Salamanca Formation

 Argentina

A member of the family Lauraceae. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.

Patagonoxylon[107]

Gen. et sp. nov

Valid

Ruiz, Brea & Pujana in Ruiz et al.

Paleocene (Danian)

Salamanca Formation

 Argentina

A Lauralean of uncertain phylogenetic placement.
The type species is P. scalariforme.
Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.

Persea masotkholaensis[103]

Sp. nov

Valid

Prasad et al.

Middle Miocene

Lower Churia Group

   Nepal

A species of Persea.

Magnoliales

Name Novelty Status Authors Age Type locality Location Notes Images

Anaxagorea mioluzonensis[103]

Sp. nov

Valid

Prasad et al.

Late Miocene

Middle Churia Group

   Nepal

A species of Anaxagorea.

Anonaspermum orientalis[108]

Sp. nov

Valid

Li et al.

Late Oligocene

Yongning Formation

 China

A member of the family Annonaceae.

Mitrephora mioreticulata[103]

Sp. nov

Valid

Prasad et al.

Late Miocene

Middle Churia Group

   Nepal

A species of Mitrephora.

Riaselis[109]

Gen. et sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous (late Aptian-early Albian or older)

 Portugal

Genus includes new species R. rugosa.

Serialis[109]

Gen. et 9 sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous (late Barremian-early Albian)

Almargem Formation
Figueira da Foz Formation

 Portugal

Genus includes new species S. antiqua, S. parva, S. elongata, S. tenuitesta, S. communis, S. crassitesta, S. grossa, S. undata and S. reticulata.

Uvaria miolucida[103]

Sp. nov

Valid

Prasad et al.

Late Miocene

Middle Churia Group

   Nepal

A species of Uvaria.

Piperales

Name Novelty Status Authors Age Type locality Location Notes Images

Appofructus[91]

Gen. et sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous (late Barremian-early Aptian)

Almargem Formation

 Portugal

Genus includes new species A. nudus.

Appomattoxia minuta[91]

Sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous (late Barremian-early Aptian)

Almargem Formation

 Portugal

Burgeria[91]

Gen. et sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous (late Barremian-early Aptian)

Almargem Formation

 Portugal

Genus includes new species B. striata.

Dejaxia[91]

Gen. et sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous (late Barremian-early Aptian)

Almargem Formation

 Portugal

Genus includes new species D. brevicolpites.

Goczania[91]

Gen. et 3 sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous (late Barremian-early Aptian)

Almargem Formation

 Portugal

Genus includes new species G. rugosa, G. inaequalis and G. punctata.

Unplaced non-eudicots

Chloranthales

Name Novelty Status Authors Age Type locality Location Notes Images

Canrightia elongata[91]

Sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous (late Barremian-early Aptian)

Almargem Formation

 Portugal

Hedyflora[110]

Gen. et sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous (late Aptian–early Albian)

Figueira da Foz Formation

 Portugal

A member of the family Chloranthaceae. Genus includes new species H. crystallifera.

Kvacekispermum costatum[91]

Sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous (late Barremian-early Aptian)

Almargem Formation

 Portugal

Basal eudicots

Proteales

Name Novelty Status Authors Age Type locality Location Notes Images

Meliosma berryi[111]

Sp. nov

Valid

Huegele & Manchester

Probably late Eocene

 United States
( Texas)

A species of Meliosma.

Platanus heilongjiangensis[112]

Sp. nov

Valid

Sun et al.

Late Cretaceous (China)

Houshiigou Formation

 China

A species of Platanus.

Scalarixylon romeroi[113]

Sp. nov

Valid

Pujana & Ruiz

Eocene–Oligocene

Río Turbio Formation

 Argentina

Ranunculales

Name Novelty Status Authors Age Type locality Location Notes Images

Tinospora siwalika[103]

Sp. nov

Valid

Prasad et al.

Late Miocene

Middle Churia Group

   Nepal

A species of Tinospora.

Superasterids

Aquifoliales

Name Novelty Status Authors Age Type locality Location Notes Images

Ilex angustifolioides[114]

Nom. nov

Valid

Doweld

Miocene

 Germany

A holly; a replacement name for Ilex denticulata von Heer (1857).

Ilex aschutassica[114]

Nom. nov

Valid

Doweld

Oligocene

 Kazakhstan

A holly; a replacement name for Ilex integrifolia Baikovskaja (1956).

Ilex boulayi[114]

Nom. nov

Valid

Doweld

Miocene

 France

A holly; a replacement name for Ilex undulata Boulay (1887).

Ilex friedrichii[114]

Nom. nov

Valid

Doweld

Oligocene

 Germany

A holly; a replacement name for Ilex longifolia Friedrich (1884).

Ilex latifolioides[114]

Nom. nov

Valid

Doweld

Oligocene

 France

A holly; a replacement name for Ilex acuminata Saporta (1865).

Ilex mormonica[114]

Nom. nov

Valid

Doweld

Oligocene

 United States
( Montana)

A holly; a replacement name for Ilex acuminata Becker (1960).

Ilex opacina[114]

Nom. nov

Valid

Doweld

Oligocene

 France

A holly; a replacement name for Ilex microdonta Saporta (1865).

Ilex polarica[114]

Nom. nov

Valid

Doweld

Paleocene

 Greenland

A holly; a replacement name for Ilex macrophylla von Heer (1869).

Ilex subrotunda[114]

Sp. nov

Valid

Doweld

Miocene

 Japan

A holly; a replacement name for the previously invalidly published Ilex ohashii Huzioka (1963), lacking holotype designation when published.

Asterales

Name Novelty Status Authors Age Type locality Location Notes Images

Cichoreacidites? igapoensis[24]

Sp. nov

Valid

D'Apolito et al.

Pliocene–Pleistocene

 Brazil

Fossil pollen of a member of the genus Pacourina or Vernonia.

Boraginales

Name Novelty Status Authors Age Type locality Location Notes Images

Cordia siwalica[103]

Sp. nov

Valid

Prasad et al.

Middle Miocene

Lower Churia Group

   Nepal

A species of Cordia.

Caryophyllales

Name Novelty Status Authors Age Type locality Location Notes Images

Basella keralensis[115]

Sp. nov

Valid

Farooqui, Ray & Garg

Pleistocene

 India

A species of Basella.

Cornales

Name Novelty Status Authors Age Type locality Location Notes Images

Eydeia jerseyensis[116]

Sp. nov

Valid

Atkinson, Martínez & Crepet

Late Cretaceous (Turonian)

 United States
( New Jersey)

Ericales

Name Novelty Status Authors Age Type locality Location Notes Images

Juddicarpon[117]

Gen. et sp. nov

Valid

Smith & Manchester

Miocene (Burdigalian-Langhian)

Clarkia fossil beds

 United States
( Idaho)

A member of Vaccinioideae. Genus includes new species J. benewahensis.

Psilastephanocolporites brevissimus[24]

Sp. nov

Valid

D'Apolito et al.

Pliocene–Pleistocene

 Brazil

Fossil pollen of a flowering plant, possibly a member of the genus Myrsine.

Sladenia zhengyii[118]

Sp. nov

Valid

Jia & Zhou in Jia et al.

Early Miocene

Maguan Basin

 China

A member of the family Sladeniaceae. Announced in 2019; the final version of the article naming it was published in 2021.

Symplocos amoena[111]

Sp. nov

Valid

Huegele & Manchester

Probably late Eocene

 United States
( Texas)

A species of Symplocos.

Symplocos fritschii[111]

Sp. nov

Valid

Huegele & Manchester

Probably late Eocene

 United States
( Texas)

A species of Symplocos.

Symplocos martinettoi[111]

Sp. nov

Valid

Huegele & Manchester

Probably late Eocene

 United States
( Texas)

A species of Symplocos.

Symplocos platycarpa[111]

Sp. nov

Valid

Huegele & Manchester

Probably late Eocene

 United States
( Texas)

A species of Symplocos.

Symplocos rothwelii[111]

Sp. nov

Valid

Huegele & Manchester

Probably late Eocene

 United States
( Texas)

A species of Symplocos.

Symplocos trinitiensis[111]

Sp. nov

Valid

Huegele & Manchester

Probably late Eocene

 United States
( Texas)

A species of Symplocos.

Symplocos trisulcata[111]

Sp. nov

Valid

Huegele & Manchester

Probably late Eocene

 United States
( Texas)

A species of Symplocos.

Gentianales

Name Novelty Status Authors Age Type locality Location Notes Images

Calycophyllum plengei[119]

Sp. nov

Valid

Woodcock, Meyer & Prado

Eocene

Piedra Chamana Fossil Forest

 Peru

A species of Calycophyllum.

Psilatriporites aspidatus[24]

Sp. nov

Valid

D'Apolito et al.

Pliocene–Pleistocene

 Brazil

Fossil pollen of a member of the genus Faramea.

Randia premacrophylla[103]

Sp. nov

Valid

Prasad et al.

Late Miocene

Middle Churia Group

   Nepal

A species of Randia.

Icacinales

Name Novelty Status Authors Age Type locality Location Notes Images

Iodes acuta[120]

Sp. nov

Valid

Del Rio, Stull & De Franceschi

Early Eocene

 France

A member of the family Icacinaceae.

Iodes parva[121]

Sp. nov

Valid

Del Rio, Thomas & De Franceschi

Late Paleocene

 France

A member of the family Icacinaceae.

Iodes reidii[121]

Sp. nov

Valid

Del Rio, Thomas & De Franceschi

Late Paleocene

 France

A member of the family Icacinaceae.

Iodes rigida[120]

Sp. nov

Valid

Del Rio, Stull & De Franceschi

Early Eocene

 France

A member of the family Icacinaceae.

Iodes rivecourtensis[121]

Sp. nov

Valid

Del Rio, Thomas & De Franceschi

Late Paleocene

 France

A member of the family Icacinaceae.

Iodes sinuosa[121]

Sp. nov

Valid

Del Rio, Thomas & De Franceschi

Late Paleocene

 France

A member of the family Icacinaceae.

Iodes tubulifera[121]

Sp. nov

Valid

Del Rio, Thomas & De Franceschi

Late Paleocene

 France

A member of the family Icacinaceae.

Superrosids

Malvids

Malvales
Name Novelty Status Authors Age Type locality Location Notes Images

Anisoptera palaeoscaphula[103]

Sp. nov

Valid

Prasad et al.

Late Miocene

Middle Churia Group

   Nepal

A species of Anisoptera.

Ceiba archeopentandra[119]

Sp. nov

Valid

Woodcock, Meyer & Prado

Eocene

Piedra Chamana Fossil Forest

 Peru

A species of Ceiba.

Ceiba huancabambiana[119]

Sp. nov

Valid

Woodcock, Meyer & Prado

Eocene

Piedra Chamana Fossil Forest

 Peru

A species of Ceiba.

Dipterocarpus palaeoindicus[103]

Sp. nov

Valid

Prasad et al.

Late Miocene

Middle Churia Group

   Nepal

A species of Dipterocarpus.

Dryobalanoxylon neosumatrense[122]

Sp. nov

Valid

Biswas, Khan & Bera

Late Miocene

 India

A member of the family Dipterocarpaceae.

Grewia americana[119]

Sp. nov

Valid

Woodcock, Meyer & Prado

Eocene

Piedra Chamana Fossil Forest

 Peru

A species of Grewia.

Grewia nepalensis[103]

Sp. nov

Valid

Prasad et al.

Middle Miocene

Lower Churia Group

   Nepal

A species of Grewia.

Grewia palaeodisperma[103]

Sp. nov

Valid

Prasad et al.

Late Miocene

Middle Churia Group

   Nepal

A species of Grewia.

Guazuma santacruzensis[119]

Sp. nov

Valid

Woodcock, Meyer & Prado

Eocene

Piedra Chamana Fossil Forest

 Peru

A member of the family Malvaceae.

Luehea stratificata[119]

Sp. nov

Valid

Woodcock, Meyer & Prado

Eocene

Piedra Chamana Fossil Forest

 Peru

A species of Luehea.

Muntingia solapora[119]

Sp. nov

Valid

Woodcock, Meyer & Prado

Eocene

Piedra Chamana Fossil Forest

 Peru

A species of Muntingia.

Ochroma pozoensis[119]

Sp. nov

Valid

Woodcock, Meyer & Prado

Eocene

Piedra Chamana Fossil Forest

 Peru

A species of Ochroma.

Sterculia arjunkholaensis[103]

Sp. nov

Valid

Prasad et al.

Middle Miocene

Lower Churia Group

   Nepal

A species of Sterculia.

Sterculia matrum[119]

Sp. nov

Valid

Woodcock, Meyer & Prado

Eocene

Piedra Chamana Fossil Forest

 Peru

A species of Sterculia.

Vasivaea weigendii[119]

Sp. nov

Valid

Woodcock, Meyer & Prado

Eocene

Piedra Chamana Fossil Forest

 Peru

Sapindales
Name Novelty Status Authors Age Type locality Location Notes Images

Ailanthus maximus[123]

Sp. nov

Valid

Liu, Su & Zhou in Liu et al.

Latest Paleocene to the Late Oligocene

Lunpola Basin
Nima Basin

 China

A species of Ailanthus.

Antrocaryon panamaensis[124]

Sp. nov

Valid

Herrera et al.

Early Miocene

Cucaracha Formation

 Panama

A species of Antrocaryon.

Arytera miolittoralis[103]

Sp. nov

Valid

Prasad et al.

Late Miocene

Middle Churia Group

   Nepal

A species of Arytera.

Arytera nepalensis[103]

Sp. nov

Valid

Prasad et al.

Late Miocene

Middle Churia Group

   Nepal

A species of Arytera.

Buchanania raptiensis[103]

Sp. nov

Valid

Prasad et al.

Late Miocene

Middle Churia Group

   Nepal

A species of Buchanania.

Dodonaea piedra-chamana[119]

Sp. nov

Valid

Woodcock, Meyer & Prado

Eocene

Piedra Chamana Fossil Forest

 Peru

A species of Dodonaea.

Dracontomelon montesii[124]

Sp. nov

Valid

Herrera et al.

Early Miocene

Cucaracha Formation

 Panama

A species of Dracontomelon.

Erythrochiton masotkholaensis[103]

Sp. nov

Valid

Prasad et al.

Late Miocene

Middle Churia Group

   Nepal

A species of Erythrochiton.

Euphoria churiaensis[103]

Sp. nov

Valid

Prasad et al.

Middle Miocene

Lower Churia Group

   Nepal

A member of the family Sapindaceae.

Koelreuteria lunpolaensis[125]

Sp. nov

Valid

Jiang et al.

Late Oligocene

Lunpola Basin

 China

A species of Koelreuteria.

Rhus asymmetrica[126]

Sp. nov

Valid

Tosal, Sanjuan & Martín-Closas

Early Oligocene

 Spain

A sumac.

Rhus boothillensis[127]

Sp. nov

Valid

Flynn, DeVore & Pigg

Early Eocene

Klondike Mountain Formation

 United States
( Washington)

A sumac.

Rhus boothillensi

Rhus garwellii[127]

Sp. nov

Valid

Flynn, DeVore & Pigg

Early Eocene

Klondike Mountain Formation

 United States
( Washington)

A sumac.

Rhus republicensis[127]

Sp. nov

Valid

Flynn, DeVore & Pigg

Early Eocene

Klondike Mountain Formation

 United States
( Washington)

A sumac.

Sapindus palaeomukorossi[103]

Sp. nov

Valid

Prasad et al.

Middle Miocene

Lower Churia Group

   Nepal

A species of Sapindus.

Spondias rothwellii[124]

Sp. nov

Valid

Herrera et al.

Early Miocene

Cucaracha Formation

 Panama

A species of Spondias.

Zanthoxylum reynelii[119]

Sp. nov

Valid

Woodcock, Meyer & Prado

Eocene

Piedra Chamana Fossil Forest

 Peru

A species of Zanthoxylum.

Other malvids
Name Novelty Status Authors Age Type locality Location Notes Images

Akania gibsonorum[128]

Sp. nov

Valid

Conran et al.

Early Miocene

 New Zealand

A member of the family Akaniaceae.

Combretum siwalicum[103]

Sp. nov

Valid

Prasad et al.

Late Miocene

Middle Churia Group

   Nepal

A species of Combretum.

Eugenia nepalensis[103]

Sp. nov

Valid

Prasad et al.

Late Miocene

Middle Churia Group

   Nepal

A species of Eugenia.

Miconioidea[119]

Gen. et sp. nov

Valid

Woodcock, Meyer & Prado

Eocene

Piedra Chamana Fossil Forest

 Peru

A member of the family Melastomataceae. Genus includes new species M. eocenica.

Myrceugenellites grandiporosum[107]

Sp. nov

Valid

Ruiz, Brea & Pujana in Ruiz et al.

Paleocene (Danian)

Salamanca Formation

 Argentina

A member of the family Myrtaceae. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.

Staphylea ochoterenae[129]

Sp. nov

Valid

Hernández-Damián, Cevallos-Ferriz & Huerta-Vergara

Miocene

 Mexico

A species of Staphylea.

Terminalia arjunkholaensis[103]

Sp. nov

Valid

Prasad et al.

Late Miocene

Middle Churia Group

   Nepal

A species of Terminalia.

Turpinia tiffneyi[111]

Sp. nov

Valid

Huegele & Manchester

Probably late Eocene

 United States
( Texas)

A species of Turpinia.

Fabids

Fabales
Name Novelty Status Authors Age Type locality Location Notes Images

Arcoa lindgreni[130]

Sp. nov

Valid

Herendeen & Herrera

Eocene

Green River Formation

 United States
( Wyoming)

A species of Arcoa.

Bauhinia palaeomonandra[103]

Sp. nov

Valid

Prasad et al.

Middle Miocene

Lower Churia Group

   Nepal

A species of Bauhinia.

Butea nepalensis[103]

Sp. nov

Valid

Prasad et al.

Late Miocene

Middle Churia Group

   Nepal

A species of Butea.

Cassia arjunkholaensis[103]

Sp. nov

Valid

Prasad et al.

Late Miocene

Middle Churia Group

   Nepal

A species of Cassia.

Cercioxylon zeynepae[43]

Sp. nov

Valid

Akkemik

Pliocene

Örencik Formation

 Turkey

A relative of redbuds described on the basis of fossil wood.

Gleditsia europaea[131]

Sp. nov

Valid

Worobiec in Worobiec & Worobiec

Miocene

 Poland

A species of Gleditsia.

Hopeoxylon umarsarensis[132]

Sp. nov

Valid

Shukla, Singh & Mehrotra

Early Eocene

Naredi Formation

 India

A member of the family Fabaceae belonging to the subfamily Detarioideae.

Leguminophyllum kvacekii[131]

Sp. nov

Valid

Worobiec in Worobiec & Worobiec

Miocene

 Poland

Fossil leaflets resembling leaflets of extant and fossil members of Fabaceae.

Millettia arjunkholaensis[103]

Sp. nov

Valid

Prasad et al.

Middle Miocene

Lower Churia Group

   Nepal

A species of Millettia.

Mimosoxylon ceratonioides[105]

Sp. nov

Valid

Akkemik in Akkemik, Akkılıç & Güngör

Early Miocene

 Turkey

Ormosia zhangpuensis[133]

Sp. nov

Valid

Wang et al.

Miocene

 China

A species of Ormosia.

Saraca palaeoindica[103]

Sp. nov

Valid

Prasad et al.

Late Miocene

Middle Churia Group

   Nepal

A species of Saraca.

Sindora eosiamensis[103]

Sp. nov

Valid

Prasad et al.

Middle Miocene

Lower Churia Group

   Nepal

A species of Sindora.

Sindora leguminocarpoides[103]

Sp. nov

Valid

Prasad et al.

Middle Miocene

Lower Churia Group

   Nepal

A species of Sindora.

Tzotziloxylon[134]

Gen. et 2 sp. nov

Valid

Pérez-Lara & Estrada-Ruiz in Pérez-Lara, Estrada-Ruiz & Castañeda-Posadas

Eocene

El Bosque Formation

 Mexico

A member of the family Fabaceae belonging to the subfamily Cercidoideae or Dialioideae. Genus includes new species T. cristalliferum and T. eocenica.

Fagales
Name Novelty Status Authors Age Type locality Location Notes Images

Castanopsis rothwellii[135][136][137]

Sp. nov

Valid

Wilf et al.

Eocene

 Argentina

A species of Castanopsis.

Casuarinoxylon ildephonsi[138]

Sp. nov

Valid

Vanner

Miocene

 New Zealand

A member of the family Casuarinaceae described on the basis of fossil wood.

Engelhardia trinitiensis[111]

Sp. nov

Valid

Huegele & Manchester

Probably late Eocene

 United States
( Texas)

A species of Engelhardia.

Pterocaryoxylon tuncayi[105]

Sp. nov

Valid

Akkemik in Akkemik, Akkılıç & Güngör

Early Miocene

 Turkey

Quercus shangcunensis[139]

Sp. nov

Valid

Liu et al.

Early Oligocene

Shangcun Formation

 China

An oak

Malpighiales
Name Novelty Status Authors Age Type locality Location Notes Images

Calophyllum mioelatum[103]

Sp. nov

Valid

Prasad et al.

Late Miocene

Middle Churia Group

   Nepal

A species of Calophyllum.

Calophyllum zhangpuensis[140]

Sp. nov

Valid

Wang et al.

Miocene

Fotan Group

 China

A species of Calophyllum.

Elioxylon[141]

Gen. et sp. nov

Valid

Srivastava, Miller & Baas

Late Cretaceous (Maastrichtian)–Paleocene (Danian)

Deccan Intertrappean Beds

 India

A wood morphospecies with features of Achariaceae and Salicaceae. Type species includes new species E. seoniensis.

Garcinia zhangpuensis[142]

Sp. nov

Valid

Wang et al.

Middle Miocene

Fotan Group

 China

A species of Garcinia.

Mascogophyllum[143]

Gen. et sp. nov

Valid

Centeno-González, Porras-Múzquiz & Estrada-Ruiz

Late Cretaceous (late Campanian)

Olmos Formation

 Mexico

A possible member of Violaceae. Genus includes new species M. elizondoa.

Populus erratica[144]

Nom. nov

Valid

Sachse

Late Oligocene and early Miocene

  Switzerland
 France?
 Germany?
 Hungary?

A species of Populus; a replacement name for Juglans heerii Ettingshausen (1853).

Ryparia arjunkholaensis[103]

Sp. nov

Valid

Prasad et al.

Late Miocene

Middle Churia Group

   Nepal

A member of the family Achariaceae.

Oxalidales
Name Novelty Status Authors Age Type locality Location Notes Images

Caldcluvioxylon torresiae[113]

Sp. nov

Valid

Pujana & Ruiz

Eocene–Oligocene

Río Turbio Formation

 Argentina

A member of the family Cunoniaceae.

Tropidogyne lobodisca[145]

Sp. nov

Valid

Poinar & Chambers

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A probable member of Cunoniaceae.

Rosales
Name Novelty Status Authors Age Type locality Location Notes Images

Artocarpus arjunkholaensis[103]

Sp. nov

Valid

Prasad et al.

Late Miocene

Middle Churia Group

   Nepal

A species of Artocarpus.

Cedrelospermum tibeticum[146]

Sp. nov

Valid

Jia, Su & Zhou in Jia et al.

Late Oligocene

Dingqing Formation

 China

A member of Ulmaceae.

Ficus preglobosa[103]

Sp. nov

Valid

Prasad et al.

Middle Miocene

Lower Churia Group

   Nepal

A species of Ficus.

Frangulops[114]

Gen. et comb. nov

Valid

Doweld

Eocene

 United States
( Colorado)

A member of Rhamnaceae; a new genus for "Ilex" pseudostenophylla Lesquereux (1883).

Prunoidoxylon aytugii[105]

Sp. nov

Valid

Akkemik in Akkemik, Akkılıç & Güngör

Early Miocene

 Turkey

Pteroceltis shanwangensis[147]

Sp. nov

Valid

Wong, Dilcher & Uemura

Miocene

Shanwang Formation

 China

A species of Pteroceltis.

Pteroceltis taoae[147]

Sp. nov

Valid

Wong, Dilcher & Uemura

Miocene

 China

A species of Pteroceltis.

Rubus eubaticus[148]

Nom. nov

Valid

Doweld

Miocene

 Bulgaria

A species of Rubus; a replacement name for Rubus mucronatus Palamarev (1987).

Rubus primoricus[148]

Nom. nov

Valid

Doweld

Miocene

 Russia
( Primorsky Krai)

A species of Rubus; a replacement name for Rubus ellipticus Pavlyutkin (2005).

Ulmus prestonia[149]

Sp. nov

Valid

Lott, Manchester & Corbett

Miocene

 United States
( Florida)

An elm.

Unplaced superrosid eudicots

Name Novelty Status Authors Age Type locality Location Notes Images

Cayratia palaeojaponica[103]

Sp. nov

Valid

Prasad et al.

Late Miocene

Middle Churia Group

   Nepal

A species of Cayratia.

Liquidambar bella[150]

Sp. nov

Valid

Maslova et al.

Eocene

Huangniuling Formation

 China

A Liquidambar species saxifragale.

Yua texana[111]

Sp. nov

Valid

Huegele & Manchester

Probably late Eocene

 United States
( Texas)

A Yua species vitale.

Other angiosperms

Name Novelty Status Authors Type locality Type locality Location Notes Images

Baccatocarpon[151]

Gen. et comb. nov

Valid

Bhowal & Sheikh ex Manchester, Ramteke, Kapgate & Smith

Late Cretaceous (Maastrichtian)

Deccan Intertrappean Beds

 India

A fossil fruit of a flowering plant of uncertain affinities; a new genus for "Grewia" mohgaoensis Paradkar & Dixit (1984).

Battenipollis sabrinae[152]

Sp. nov

Valid

Smith et al.

Early Paleogene

Antarctica

An angiosperm pollen species.

Bonanzacarpum[153]

Gen. et sp. nov

Valid

Manchester & Lott

Early to middle Eocene

Green River Formation

 United States
( Utah)

A eudicot fossil fruit of uncertain phylogenetic placement.
The type species is B. sprungerorum.

Celastrilex[114]

Gen. et comb. nov

Valid

Doweld

Paleocene

 United States
( Colorado)

A flowering plant of uncertain phylogenetic placement, described on the basis of fossil leaves; a new genus for "Celastrinites" artocarpidioides Lesquereux (1878).

Choffaticarpus[91]

Gen. et sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous (late Barremian-early Aptian)

Almargem Formation

 Portugal

A flowering plant of uncertain phylogenetic placement. Genus includes new species C. compactus.

Cratolirion[154]

Gen. et sp. nov

Valid

Coiffard, Kardjilov et Bernardes-de-Oliveira in Coiffard et al.

Early Cretaceous

Crato Formation

 Brazil

A crown monocot of uncertain phylogenetic placement. Genus includes new species C. bognerianum.

Dalembia (?) gracilis[155]

Sp. nov

Valid

Herman in Herman et al.

Late Cretaceous (Turonian-Coniacian)

Derevyannye Gory Formation

 Russia
( Sakha Republic)

A flowering plant described on the basis of fossil leaves.

Dictyozonia[91]

Gen. et sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous (late Barremian-early Aptian)

Almargem Formation

 Portugal

A non-eudicot flowering plant of uncertain affinity. Genus includes new species D. pusilla.

Dinisia[91]

Gen. et sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous (late Barremian-early Aptian)

Almargem Formation

 Portugal

A non-eudicot flowering plant of uncertain affinity. Genus includes new species D. portugallica.

Dispariflora[156]

Gen. et sp. nov

Valid

Poinar & Chambers

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A flowering plant of uncertain phylogenetic placement, possibly a relative of members of Laurales, especially Southern Hemisphere families allied with the Monimiaceae. Genus includes new species D. robertae.

Eckhartia[91]

Gen. et 3 sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous (late Barremian-early Aptian)

Almargem Formation

 Portugal

A non-eudicot flowering plant of uncertain affinity. Genus includes new species E. brevicolumella, E. longicolumella and E. intermedia.

Eckhartianthus[91]

Gen. et sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous (late Barremian-early Aptian)

Almargem Formation

 Portugal

A non-eudicot flowering plant of uncertain affinity. Genus includes new species E. lusitanicus.

Eckhartiopsis[91]

Gen. et sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous (late Barremian-early Aptian)

Almargem Formation

 Portugal

A non-eudicot flowering plant of uncertain affinity. Genus includes new species E. parva.

Exalloanthum[157]

Nom. nov

Valid

Poinar

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A flowering plant of uncertain phylogenetic placement; a replacement name for Diaphoranthus Poinar (2018).

Gambierina askiniae[152]

Sp. nov

Valid

Smith et al.

Early Paleogene

Antarctica

An angiosperm pollen species.

Herendeenoxylon[158]

Gen. et sp. nov

Valid

Chin et al.

Late Cretaceous (Turonian)

Moreno Hill Formation

 United States
( New Mexico)

A flowering plant of uncertain phylogenetic placement (possibly a member of Ericales), described on the basis of fossil wood. Genus includes new species H. zuniense.

Ibrahimia[91]

Gen. et sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous (late Barremian-early Aptian)

Almargem Formation

 Portugal

An eudicot of uncertain phylogenetic placement, possibly related to Paisia. Genus includes new species I. vermiculata.

Juhaszia[91]

Gen. et sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous (late Barremian-early Aptian)

Almargem Formation

 Portugal

A non-eudicot flowering plant of uncertain affinity. Genus includes new species J. portugallica.

Kempia[91]

Gen. et sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous (late Barremian-early Aptian)

Almargem Formation

 Portugal

A non-eudicot flowering plant of uncertain affinity. Genus includes new species K. longicolpites.

Ladakhipollenites? densicolumellatus[24]

Sp. nov

Valid

D'Apolito et al.

Pliocene–Pleistocene

 Brazil

Fossil pollen of a flowering plant.

Ladakhipollenites? lolongatus[24]

Sp. nov

Valid

D'Apolito et al.

Pliocene–Pleistocene

 Brazil

Fossil pollen of Symmeria paniculata.

Ladakhipollenites? porolenticularis[24]

Sp. nov

Valid

D'Apolito et al.

Pliocene–Pleistocene

 Brazil

Fossil pollen of a flowering plant (possibly a member of the family Marcgraviaceae).

Lagokarpos tibetensis[159]

Sp. nov

Valid

Tang, Su & Zhou in Tang et al.

Paleogene

Niubao Formation

 China

A fossil fruit with unknown modern affinities.

Mcdougallia[91]

Gen. et sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous (late Barremian-early Aptian)

Almargem Formation

 Portugal

An eudicot of uncertain phylogenetic placement. Genus includes new species M. irregularis.

Nicholsia[91]

Gen. et sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous (late Barremian-early Aptian)

Almargem Formation

 Portugal

An eudicot of uncertain phylogenetic placement. Genus includes new species N. brevicolpites.

Piercipollis[91]

Gen. et sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous (late Barremian-early Aptian)

Almargem Formation

 Portugal

A non-eudicot flowering plant of uncertain affinity. Genus includes new species P. simplex.

Reyanthus[91]

Gen. et sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous (late Barremian-early Aptian)

Almargem Formation

 Portugal

A flowering plant of uncertain phylogenetic placement, possibly related to Magnoliales. Genus includes new species R. lusitanicus.

Rhoipites? basicus[24]

Sp. nov

Valid

D'Apolito et al.

Pliocene–Pleistocene

 Brazil

Fossil pollen of a flowering plant.

Rhoipites manausensis[24]

Sp. nov

Valid

D'Apolito et al.

Pliocene–Pleistocene

 Brazil

Fossil pollen of a member of the genus Schefflera.

Rhoipites minuticirculus[24]

Sp. nov

Valid

D'Apolito et al.

Pliocene–Pleistocene

 Brazil

Fossil pollen of a flowering plant.

Rhoipites negroensis[24]

Sp. nov

Valid

D'Apolito et al.

Pliocene–Pleistocene

 Brazil

Fossil pollen of a flowering plant.

Samylinaea[91]

Gen. et sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous (late Barremian-early Aptian)

Almargem Formation

 Portugal

An eudicot of uncertain phylogenetic placement. Genus includes new species S. punctata.

Sherwinoxylon[160]

Gen. et sp. nov

Valid

Boura & Saulnier in Boura et al.

Late Cretaceous (Cenomanian)

 France

A vesselless angiosperm fossil wood of uncertain affinity. Genus includes new species S. winteroides.

Strombothelya[161]

Gen. et 2 sp. nov

Valid

Poinar & Chambers

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A flowering plant of uncertain phylogenetic placement. Genus includes new species S. monostyla and S. grammogyna.

Teebacia[91]

Gen. et sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous (late Barremian-early Aptian)

Almargem Formation

 Portugal

A non-eudicot flowering plant of uncertain affinity. Genus includes new species T. hughesii.

Ubiquitoxylon[162]

Gen. et sp. nov

Valid

Wheeler in Wheeler, Brown & Koch

Late Paleocene

Denver Formation

 United States
( Colorado)

A dicotyledonous flowering plant of uncertain phylogenetic placement, described on the basis of fossil wood. Genus includes new species U. raynoldsii.

Vasunum[158]

Gen. et sp. nov

Valid

Chin et al.

Late Cretaceous (Turonian)

Moreno Hill Formation

 United States
( New Mexico)

A flowering plant of uncertain phylogenetic placement, described on the basis of fossil wood. Genus includes new species V. cretaceum.

Vedresia[91]

Gen. et sp. nov

Valid

Friis, Crane & Pedersen

Early Cretaceous (late Barremian-early Aptian)

Almargem Formation

 Portugal

A flowering plant of uncertain phylogenetic placement, possibly related to Chloranthales. Genus includes new species V. elliptica.

Zygadelphus[163]

Gen. et sp. nov

Valid

Poinar & Chambers

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A flowering plant of uncertain phylogenetic placement, possibly a member of Laurales. Genus includes new species Z. aetheus.

Other plants

Name Novelty Status Authors Age Type locality Location Notes Images

Acetabularia moldavica[164]

Sp. nov

Valid

Barattolo, Ionesi & Ţibuleac

Middle Miocene

 Romania

A green alga belonging to the family Polyphysaceae, a species of Acetabularia.

Aloisalthella[165]

Gen. et comb. nov

Valid

Granier in Granier & Lethiers

Late Jurassic and Early Cretaceous (Berriasian)

 Algeria
 France
 Spain
 Ukraine
 United Arab Emirates

A green alga belonging to the family Polyphysaceae; a new genus for "Actinoporella" sulcata von Alth (1882).

Aneurospora posongchongensis[166]

Sp. nov

Valid

Cascales-Miñana et al.

Early Devonian

Posongchong Formation

 China

A spore taxon.

Auerbachichara tataouinensis[167]

Sp. nov

Valid

Tiss et al.

Middle Jurassic (Callovian)

Foum Tataouine Formation

 Tunisia

A green alga belonging to the group Charophyta.

Bakalovaella deloffrei[168]

Sp. nov

Valid

Granier & Bucur

Early Cretaceous (Hauterivian)

 France

A green alga belonging to the family Dasycladaceae.

Buthograptus gundersoni[169]

Sp. nov

Valid

LoDuca

Ordovician (Sandbian)

Platteville Formation

 United States
( Wisconsin)

A green alga belonging to the group Bryopsidales.

Buthograptus meyeri[169]

Sp. nov

Valid

LoDuca

Ordovician (Sandbian)

Platteville Formation

 United States
( Wisconsin)

A green alga belonging to the group Bryopsidales.

Calcipatera schoenlaubi[170]

Sp. nov

Valid

Vachard in Krainer, Vachard & Schaffhauser

Permian

 Austria
 Oman?
 United States?
( New Mexico)

A green alga belonging to the group Bryopsidales and the family Anchicodiaceae.

Callixylon wendtii[171]

Sp. nov

Valid

Tanrattana, Meyer-Berthaud & Decombeix

Devonian (Famennian)

 Morocco

An archaeopteridalean progymnosperm.

Cingulatisporites oligodistalis[24]

Sp. nov

Valid

D'Apolito et al.

Pliocene–Pleistocene

 Brazil

Fossil spore.

Concavissimisporites varzeanus[24]

Sp. nov

Valid

D'Apolito et al.

Pliocene–Pleistocene

 Brazil

Fossil spore.

Coniopteris moguqiensis[172]

Sp. nov

Valid

Zhang, Liu & Liang

Middle Jurassic

Wanbao Formation

 China

Dissocladella? chahtorshiana[173]

Sp. nov

Valid

Rashidi & Schlagintweit in Schlagintweit et al.

Paleocene

 Iran

A green alga belonging to the family Dasycladaceae.

Dissocladella compressa[174]

Sp. nov

Valid

Rashidi & Schlagintweit

Late Cretaceous (Maastrichtian)

Tarbur Formation

 Iran

A green alga belonging to the group Dasycladales.

Echinatisporis parviechinatus[24]

Sp. nov

Valid

D'Apolito et al.

Pliocene–Pleistocene

 Brazil

Fossil spore.

Echinosporis conicus[24]

Sp. nov

Valid

D'Apolito et al.

Pliocene–Pleistocene

 Brazil

Fossil spore.

Electrophycus[175]

Gen. et sp. nov

Valid

Poinar & Brown

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A green alga, possibly a member of the family Chaetophoraceae. Genus includes new species E. astroplethus. Announced in 2019; the final version of the article naming it was published in 2021.

Epiastopora[170]

Gen. et comb. nov

Valid

Vachard in Krainer, Vachard & Schaffhauser

Carboniferous (Pennsylvanian) and Permian

A green alga belonging to the group Dasycladales and the family Seletonellaceae. A new genus for "Epimastopora" alpina Kochansky & Herak (1960) and several other species formerly assigned to the genera Epimastopora and Pseudoepimastopora.

Jowingera[176]

Gen. et sp. nov

Valid

Bickner & Tomescu

Devonian (Emsian)

Battery Point Formation

 Canada
( Quebec)

An early euphyllophyte. Genus includes new species J. triloba.

Leonophyllum[177]

Gen. et sp. nov

Valid

Barbacka & Kustatscher in Barbacka et al.

Early Jurassic

 Hungary

A plant of uncertain phylogenetic placement, showing similarities to thalloid liverworts with raised vegetative bodies and to the fern family Hymenophyllaceae.
Genus includes new species L. tenellum.

Leptocentroxyla[176]

Gen. et sp. nov

Valid

Bickner & Tomescu

Devonian (Emsian)

Battery Point Formation

 Canada
( Quebec)

An early euphyllophyte. Genus includes new species L. tetrarcha.

Maiaspora[178]

Gen. et sp. nov

Valid

Mamontov et al.

Carboniferous (Viséan)

Moscow Syneclise

 Russia

A miospore. Genus includes new species M. panopta. Announced in 2019; the final version of the article naming it was published in 2021.

Ninsaria[179]

Gen. et sp. nov

Valid

Decombeix, Galtier, McLoughlin & Meyer-Berthaud in Decombeix et al.

Carboniferous (Viséan)

Rockhampton Group

 Australia

A vascular plant belonging to the group Lignophytia, of uncertain phylogenetic placement within the latter group. Genus includes new species N. australiana.

Palambages pariunta[180]

Sp. nov

Valid

Wainman et al.

Late Jurassic (late Kimmeridgian–early Tithonian)

Surat Basin

 Australia

A colonial alga belonging to the group Chlorophyta.

Patruliuspora[164]

Gen. et comb. nov

Valid

Barattolo, Ionesi & Ţibuleac

Late Triassic to Miocene

 Czech Republic
 France
 Slovakia

A green alga belonging to the family Polyphysaceae. Genus includes "Chalmasia" morelleti Pokorný (1948), "Halicoryne" carpatica Mišík (1987) and "Acicularia" valeti Segonzac (1970).

Porochara schudackii[167]

Sp. nov

Valid

Tiss et al.

Middle Jurassic (Bajocian)

Krachoua Formation

 Tunisia

A green alga belonging to the group Charophyta.

Pseudocymopolia acuta[181]

Sp. nov

Valid

Rashidi & Schlagintweit

Late Cretaceous (Maastrichtian)

Tarbur Formation

 Iran

A green alga belonging to the group Dasycladales and to the family Triploporellaceae.

Stenoloboxyla[176]

Gen. et sp. nov

Valid

Bickner & Tomescu

Devonian (Emsian)

Battery Point Formation

 Canada
( Quebec)

An early euphyllophyte. Genus includes new species S. ambigua.

Tainioxyla[176]

Gen. et sp. nov

Valid

Bickner & Tomescu

Devonian (Emsian)

Battery Point Formation

 Canada
( Quebec)

An early euphyllophyte. Genus includes new species T. quebecana.

Tichavekia[182]

Gen. et sp. nov

Valid

Pšenička, Sakala & Kraft in Kraft et al.

Late Silurian

Prague Basin

 Czech Republic

A large early land plant. Genus includes new species T. grandis.

Uteria naghanensis[183]

Sp. nov

Valid

Rashidi & Schlagintweit

Late Cretaceous (Maastrichtian)

Tarbur Formation

 Iran

A green alga belonging to the family Polyphysaceae.

Verrucatotriletes laesuraverrucatus[24]

Sp. nov

Valid

D'Apolito et al.

Pliocene–Pleistocene

 Brazil

Fossil spore.

Verrucatotriletes tortus[24]

Sp. nov

Valid

D'Apolito et al.

Pliocene–Pleistocene

 Brazil

Fossil spore.

General research

  • Description of fossils of filamentous green algae from the Early Devonian Rhynie chert (Scotland) is published by Wellman, Graham & Lewis (2019).[184]
  • Cretaceous alga Falsolikanella campanensis, originally assigned to the tribe Diploporeae within the green alga order Dasycladales, is transferred to the genus Actinoporella within the tribe Acetabularieae, family Polyphysaceae by Barattolo et al. (2019).[185]
  • A study on the impact of the Cretaceous–Paleogene extinction event on European charophytes is published by Vicente, Csiki-Sava & Martín-Closas (2019).[186]
  • The oldest known trilete spore assemblages reported so far are described from the Sandbian successions from Motala (central Sweden) by Rubinstein & Vajda (2019).[187]
  • A study on the composition and distribution of dispersed spore assemblages from Middle Devonian deposits of northern Spain, and on their implications for inferring the nature of the Kačák Event, is published by Askew & Wellman (2019).[188]
  • A study on the morphology of the spore taxon Lagenoisporites magnus from the Carboniferous (Tournaisian) Toregua Formation (Bolivia) is published by Quetglas, Macluf & di Pasquo (2019).[189]
  • A review of research concerning early evolution of land plants during the Ordovician is published by Servais et al. (2019).[190]
  • A study on carbon isotope data from stratigraphic sections at Germany Valley (West Virginia) and Union Furnace (Pennsylvania) in the Central Appalachian Basin, evaluating its implications for the knowledge of change in atmospheric oxygen levels during the late Ordovician and its possible relationship with early diversification of land plants, is published by Adiatma et al. (2019).[191]
  • A study on the stable carbon isotopic composition of 190 fossil specimens belonging to 12 genera of Devonian and Early Carboniferous land plants is published by Wan et al. (2019).[192]
  • A study on the early evolution of vascular plants is published by Cascales‐Miñana et al. (2019).[193]
  • A study on the evolution of early vascular plants is published by Crepet & Niklas (2019).[194]
  • A study on the fine‐scale structure and the chemistry of the tracheids of the earliest known woody plant Armoricaphyton chateaupannense is published by Strullu‐Derrien et al. (2019).[195]
  • A study on diversity and functions of lycopsid reproductive structures through time, based on data from extant and fossil taxa, is published by Bonacorsi & Leslie (2019).[196]
  • Redescription of the morphology of sterile and fertile structures of the Devonian lycopsid Kossoviella timanica is published by Orlova et al. (2019).[197]
  • A study on the ultrastructure of the spore wall in the Carboniferous lycopsid Oxroadia gracilis is published by Taylor (2019).[198]
  • A slab containing rooting systems which probably belonged to rhizomorphic lycopsids is reported from the Lower Permian Abo Formation (New Mexico, United States) by Hetherington et al. (2019).[199]
  • A study on the anatomy and affinities of Cheirostrobus pettycurensis is published by Neregato & Hilton (2019), who report the discovery of spores conforming to the species Retusotriletes incohatus associated with fossils of Cheirostrobus, representing the first discovery of Retusotriletes-type spores reported in situ within sphenophytes.[200]
  • A study on the anatomy and affinities of silicified stems of Sphenophyllum from the Tournaisian deposits in the Montagne Noire region of France and in the Saalfeld area in Germany is published by Terreaux de Felice, Decombeix & Galtier (2019).[201]
  • Fossils assigned to the genus Equisetum are reported from a new fossil plant assemblage of late Eocene or early Oligocene age from central Queensland (Australia) by Rozefelds et al. (2019), representing the first evidence of this genus from the Cenozoic of Australia and the most recent fossil record of this genus from Australia.[202]
  • A study on the evolutionary history of horsetails, based on genetic data and fossil record, is published by Clark, Puttick & Donoghue (2019), who report evidence indicative of two successive whole-genome duplication events occurring during the Carboniferous and Triassic rather than in association with the Cretaceous–Paleogene extinction event.[203]
  • A study aiming to determine links between volcanic activity in the Central Atlantic magmatic province, elevated concentrations of mercury in marine and terrestrial sediments and abnormalities of fossil fern spores across the Triassic-Jurassic boundary in southern Scandinavia and northern Germany is published by Lindström et al. (2019).[204]
  • A study on the fossil record of fern spores at the Cretaceous-Paleogene boundary, on the viability of fern spores, and on their implications for the knowledge of the duration of the impact winter at the Cretaceous-Paleogene boundary is published by Berry (2019).[205]
  • A study on the molecular structural characteristics of organic remains of a fern belonging to the family Osmundaceae from the Early Jurassic Korsaröd site in southern Sweden is published by Qu et al. (2019).[206]
  • A study on anatomy and growth of large specimens of the fossil fern species Weichselia reticulata from the Barremian La Huérguina Formation (Spain) is published by Blanco-Moreno et al. (2019).[207]
  • A study on the morphological characters of 42 fossil species of Dicksoniaceae from China, and on their implications for the taxonomy of the fossil members of this group, is published by Xin et al. (2019).[208]
  • Fossil occurrences of members of the genus Christella are reported from the late Paleocene of Liuqu, southern Tibet and middle Miocene of the Jinggu Basin in western Yunnan (China) by Xu et al. (2019), who transfer the species "Cyclosorus" nervosus Tao (1988) to the genus Christella.[209]
  • A study on the fossils of Glossopteris from the Permian succession of eastern India, aiming to identify the molecular signatures of solvent-extractable and non-extractable organic matter, will be published by Tewari et al. (2019).[210]
  • A study on the diversity trends of Glossopteris flora from the Barakar, Raniganj, and Panchet formations of Tatapani–Ramkola Coalfield (India) is published by Saxena et al. (2019).[211]
  • A study on the architecture of the ovuliferous reproductive organs of Permian glossopterids is published by Mcloughlin & Prevec (2019).[212]
  • A study on the pinnule and stomatal morphology of extant and fossil members of the genera Bowenia and Eobowenia, and on its implications for the knowledge of adaptations of fossil plants to different environments, is published by Hill, Hill & Watling (2019).[213]
  • Seed of the ginkgoalean Yimaia capituliformis with damage interpreted as likely oviposition lesions inflicted by a kalligrammatid lacewing is described from the Middle Jurassic Jiulongshan Formation (China) by Meng et al. (2019).[214]
  • A study on the phytogeographic history of ten conifer genera that are endemic to East Asia, based on fossil data from humid temperate forests in the Japanese Islands and Korean Peninsula, is published by Yabe et al. (2019).[215]
  • A study on the evolution of male and female cone sizes in members of the family Araucariaceae, as indicated by data from extant and fossil members of this family, is published by Gleiser et al. (2019).[216]
  • Five fossil foliage specimens of Calocedrus lantenoisi, representing one of the earliest records of the genus Calocedrus worldwide, are described from the Oligocene Shangcun Formation of the Maoming Basin (Guangdong Province, South China) by Wu et al. (2019).[217]
  • Leaves including cuticles and ovuliferous cones of members of the genus Metasequoia are described from the middle Miocene of Zhenyuan, Yunnan (Southwest China) by Wang et al. (2019), comprising the southernmost fossil record of this genus worldwide.[218]
  • A review of the fossil record of woods which might have affinities with Taxaceae, and a study on the palaeobiogeographical history of this family, is published by Philippe et al. (2019).[219]
  • Putative Cretaceous siliceous sponge Siphonia bovista is reinterpreted as an internal mould of the cone-like plant fossil Dammarites albens by Niebuhr (2019).[220]
  • A review of epidermal features of bennettites, comparing them with analogous features in living taxa and aiming to identify homologous character states, is published by Rudall & Bateman (2019).[221]
  • The first fossil record of a cycad seedling found in close association with a leaf flush of an adult cycad plant of the same species (Dioonopsis praespinulosa) is reported from the Palaeocene (Danian) Castle Rock flora in the Denver Basin (Colorado, United States) by Erdei et al. (2019).[222]
  • A review of the paleobotanical evidence of the age and early history of the flowering plants is published by Coiro, Doyle & Hilton (2019).[223]
  • A study aiming to establish when the flowering plants originated is published by Li et al. (2019).[224]
  • Presence of endothelium (a specialized seed tissue that develops from the inner epidermis of the inner integument) is reported in several different kinds of flowering plant seeds (including in the lineage leading to extant Chloranthaceae) from the Early Cretaceous of eastern North America and Portugal by Friis, Crane & Pedersen (2019).[225]
  • A study on the phylogenetic relationships of palm fruit fossils from the Cretaceous–Paleogene (MaastrichtianDanian) Deccan Intertrappean Beds (India) is published by Matsunaga et al. (2019), who interpret these fossils as representing a crown group member of palm subtribe Hyphaeninae (tribe Borasseae, subfamily Coryphoideae) related to extant genera Satranala and Bismarckia.[226]
  • Fossil fruits of members of the genera Fragaria and Rubus are reported from the Pliocene outcrops in the Heqing Basin (China) by Huang et al. (2019).[227]
  • Description of alder leaf and infructescence fossils from the Upper Eocene Lawula Formation (Qinghai–Tibetan Plateau) is published by Xu, Su & Zhou (2019).[228]
  • A study on the morphology, paleoecology, historical biogeography and phylogenetic relationships of fossil pollen of members of Malvaceae belonging to the species Rhoipites guianensis and Malvacipolloides maristellae, and on its implications for inferring the impact of Cenozoic geological processes (including the uplift of the Andes) on members of Malvaceae living in northern South America, is published by Hoorn et al. (2019).[229]
  • A study aiming to determine the location of refugia of two North American species of hickories during the Last Glacial Maximum on the basis of genomic data is published by Bemmels, Knowles & Dick (2019).[230]
  • A study on functional leaf traits of the Eocene-Miocene taxa Rhodomyrtophyllum reticulosum (family Myrtaceae) and Platanus neptuni (family Platanaceae), evaluating whether leaf traits of these taxa reflect environmental conditions including climate, is published by Moraweck et al. (2019).[231]
  • A study on the morphology and phylogenetic relationships of Eocene fruits belonging to the species Mastixicarpum crassum and Eomastixia bilocularis is published by Manchester & Collinson (2019).[232]
  • Seeds of Eurya stigmosa are reported from the Early Pleistocene lacustrine and fluvial sediments of Porto da Cruz, Madeira by Góis-Marques et al. (2019).[233]
  • A study on the putative cycad "Zamia" australis from the Miocene Ñirihuau Formation (Argentina) is published by Passalia, Caviglia & Vera (2019), who reinterpret the fossil specimens as flowering plant leaves, and transfer this species to the genus Lithraea.[234]
  • New method for reconstructing water transport properties of fossil wood is proposed by Tanrattana et al. (2019).[235]
  • Signatures of Devonian (Famennian) forests and soils preserved in black shales in the southernmost Appalachian Basin (Chattanooga Shale; Alabama, United States) are presented by Lu et al. (2019).[236]
  • A study on reproductive structures of Devonian plants and on their implications for the knowledge of large-scale patterns of reproductive evolution over the Devonian is published by Bonacorsi & Leslie (2019).[237]
  • Revision of a fossil plant assemblage from the Carboniferous site in San Juan Province, Argentina known as Retamito or Río del Agua is published by Correa & Césari (2019).[238]
  • A study on the stratigraphic ranges and diversities of plant taxa from the upper Permian (Lopingian) to the Middle Triassic is published by Nowak, Schneebeli-Hermann & Kustatscher (2019), who interpret their findings as indicating that the extinction of land plants during the Permian–Triassic extinction event was much less severe than previously thought.[239]
  • A study on the timing of the collapse of the Permian Glossopteris flora from the Sydney Basin (Australia) is published by Fielding et al. (2019).[240]
  • New fossil flora dominated by cuticles of Dicroidium is reported from the Middle Triassic (Anisian) Mukheiris Formation (Jordan) by Abu Hamad et al. (2019).[241]
  • A study on changes of land vegetation resulting from the Toarcian oceanic anoxic event is published by Slater et al. (2019).[242]
  • Plant disseminules are documented from four Middle Jurassic to Lower Cretaceous lacustrine Lagerstätten in China and Australia by McLoughlin & Pott (2019).[243]
  • A study comparing the Jurassic floras of the Ayuquila Basin and the Otlaltepec Basin (Mexico) and evaluating their implications for the knowledge of the Jurassic environments of these basins is published by Velasco-de León et al. (2019).[244]
  • A study on phototropism in extant trees from Beijing and Jilin Provinces and fossil tree trunks from the Jurassic Tiaojishan and Tuchengzi formations in Liaoning and Beijing regions (China), and on its implications for inferring the history of the rotation of the North China Block, is published by Jiang et al. (2019).[245]
  • A study on the link between climatic changes and changes plant distribution in South America during the Early Cretaceous, as indicated by palynological data from the Aptian of the Sergipe Basin (Brazil), is published by Carvalho et al. (2019).[246]
  • A study on the frequency and diversity of damage types caused by insect oviposition in plants from the Upper Triassic Yangcaogou Formation, Middle Jurassic Jiulongshan Formation and Lower Cretaceous Yixian Formation (China), assessing the degree of plant host specificity, is published by Lin et al. (2019).[247]
  • A study on the plant specimens (ferns, gymnosperms and angiosperms) from the Lower Cretaceous Araripe Basin (Brazil) preserving evidence of plant–insect interactions and potentially of paleoecological relationships between plants and insects is published by Edilson Bezerra dos Santos Filho et al. (2019).[248]
  • Leaves of members of the family Nymphaeaceae preserving evidence of insect herbivory are reported from the Albian Utrillas Formation (Spain) by Estévez-Gallardo et al. (2019).[249]
  • A study on Cenomanian plants from the Redmond no.1 mine near Schefferville (Redmond Formation; Labrador Peninsula, Canada) and on their implications for the knowledge of paleoclimate of this site is published by Demers‐Potvin & Larsson (2019).[250]
  • A study on the canopy structure of Late Cretaceous and Paleocene forests in South America, as indicated by the carbon isotope composition of fossil angiosperm leaves from two localities in the Paleocene Cerrejón Formation and one locality in the Maastrichtian Guaduas Formation (Colombia), is published by Graham et al. (2019).[251]
  • A quantitative analysis of an earliest Paleocene megaflora from the Ojo Alamo Sandstone in the San Juan Basin (New Mexico, United States) is published by Flynn & Peppe (2019).[252]
  • A study on the evolution of plant assemblages in the area of Primorye (Russia) throughout the Paleogene is published by Bondarenko, Blokhina & Utescher (2019).[253]
  • A study on changes in plant and insect communities across the PaleoceneEocene boundary within the Hanna Basin (Wyoming, United States) is published by Azevedo Schmidt et al. (2019).[254]
  • A study on stomata of fossil specimens of members of the family Lauraceae from the Eocene of Australia and New Zealand, evaluating their implications for reconstructions of Eocene pCO2 levels, is published by Steinthorsdottir et al. (2019).[255]
  • Description of early Eocene leaf fossils from the Dinmore locality (Redbank Plains Formation, Booval Basin; Australia) and a study on the implications of these fossils for reconstructions of paleoclimate is published by Pole (2019).[256]
  • A study on changes of plant communities from the Herren beds (Oregon, United States) during the Eocene and on the implications of plant fossils from this area for the reconstruction of Eocene climate is published by Jijina, Currano & Constenius (2019).[257]
  • Su et al. (2019) use radiometrically dated plant fossil assemblages to quantify when southeastern Tibet achieved its present elevation, and what kind of floras existed there at that time.[258]
  • Description of a plant megafossil assemblage from the Kailas Formation in western part of the southern Lhasa terrane, and a study on its implications for inferring the elevation history of the southern Tibetan Plateau, is published by Ai et al. (2019).[259]
  • A study on the dynamics and evolution of the flora of Turgai ecological type in Western Siberia during the early Oligocene to earliest Miocene is published by Popova et al. (2019).[260]
  • A study on the paleoclimate, vegetational type and ecological strategies adopted by fossil plants from the Oligocene Baigang Formation (China), as indicated by characteristics of fossil leaves from this formation, is published by Li et al. (2019).[261]
  • Description of a fossil plant assemblage from the Miocene Hattiesburg Formation (Mississippi, United States) is published by McNair et al. (2019).[262]
  • A study on changes of C4 vegetation composition in southwestern Montana (United States) from the late Miocene through present is published by Hyland et al. (2019).[263]
  • A study aiming to test the hypothesis that fire contributed to the rise of C3-dominated grasslands in Eurasia, based on data from core retrieved from the late Miocene to Pleistocene sediments from the Black Sea, is published by Feurdean & Vasiliev (2019).[264]
  • A study on the origin of the African C4 savannah grasslands is published by Polissar et al. (2019).[265]
  • A study on vegetation changes in west African tropical montane forest over the past 90,000 years, as indicated by pollen data from the Lake Bambili site (Cameroon), is published by Lézine et al. (2019).[266]
  • A study on changes of vegetation in southern Borneo over the past 40,000  calibrated years BP, as indicated by data from Saleh Cave (South Kalimantan, Indonesia), is published by Wurster et al. (2019).[267]
  • A study on the role of past climate, extinct megafauna and guanaco in shaping the vegetation of the Patagonian steppe is published by Hernández, Ríos & Perotto-Baldivieso (2019).[268]
  • The discovery of ancient chestnut, hazelnut and flax DNA recovered from stalagmites from the Solkota cave (Georgia) is reported by Stahlschmidt et al. (2019).[269]
  • The discovery of oldest fossil trees, dating back 386 million years, in the Catskill region near Cairo, New York, is published online by Stein et al. (2019).[270]

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