Gymnotrachelus
Temporal range: Late Devonian: Famennian,
Artist's reconstruction
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Placodermi
Order: Arthrodira
Suborder: Brachythoraci
Family: Selenosteidae
Genus: Gymnotrachelus
Dunkel & Bungart, 1939
Species
  • Gymnotrachelus hydei Dunkel & Bungart, 1939 (type)

Gymnotrachelus is an extinct monospecific genus of large selenosteid arthrodire placoderm of the Late Devonian known from the Late Famennian Cleveland Shale of Ohio.[1] The type species Gymnotrachelus hydei was originally reconstructed as physically resembling Selenosteus, with slightly smaller orbits (i.e., having a broad, box-like head). Later specimens led to a reappraisal, and now G. hydei is thought to have a more gar-like or barracuda-like build.[2]

Phylogeny

Gymnotrachelus is a member of the family Selenosteidae of the clade Aspinothoracidi, which belongs to the clade Pachyosteomorphi, one of the two major clades within Eubrachythoraci. The cladogram below shows the phylogeny of Gymnotrachelus:[3]

Eubrachythoraci 

Coccosteomorphi

 Pachyosteomorphi 

Rhachiosteus pterygiatus

Dunkleosteoidea 

Eastmanosteus calliaspis

Eastmanosteus pustulosus

Kiangyousteus yohii

Golshanichthys asiatica

Westralichthys uwagedensis

Dunkleosteus raveri

Dunkleosteus terrelli

Dunkleosteus amblyodoratus

 Heterostiidae 

Heterosteus ingens

Yinosteus major

Aspinothoracidi

Tapinosteus heintzi

Bullerichthys fascidens

Kendrickichthys cavernosus

Bruntonichthys multidens

Dinichthys herzeri

Hadrosteus rapax

Gorgonichthys clarki

 Selenosteidae 

Heintzichthys gouldii

Pachyosteus bulla

Gymnotrachelus hydei

Stenosteus angustopectus

Brachyosteus dietrichi

Melanosteus occitanus

Rhinosteus parvulus

According to a 2022 Jobbins et al. study, Gymnotrachelus was found to be an outgroup to Heintzichthys and Gorgonichthys.[4]

Gymnotrachelus

Gorgonichthys

Heintzichthys

References

  1. Denison, Robert (1978). Handbook of Paleoichthyology, Volume 2, Placodermi. New York: Gustav Fischer Verlage. p. 96. ISBN 9780895740274.
  2. Carr, Robert (1994). "A redescription of Gymnotrachelus hydei (Placodermi : Arthrodira) from the Cleveland Shale (Famennian) of north2 ern Ohio, U. S. A." (PDF). Kirtlandia (48): 3–21.
  3. Zhu, You-An; Zhu, Min; Wang, Jun-Qing (1 April 2016). "Redescription of Yinostius major (Arthrodira: Heterostiidae) from the Lower Devonian of China, and the interrelationships of Brachythoraci". Zoological Journal of the Linnean Society. 176 (4): 806–834. doi:10.1111/zoj.12356. ISSN 0024-4082.
  4. Jobbins, Melina; Rücklin, Martin; Ferrón, Humberto G.; Klug, Christian (2022). "A new selenosteid placoderm from the Late Devonian of the eastern Anti-Atlas (Morocco) with preserved body outline and its ecomorphology". Frontiers in Ecology and Evolution. 10. doi:10.3389/fevo.2022.969158. ISSN 2296-701X.


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