Taxonomy of Lucanidae

The Lucanidae are a family of beetles that include the stag beetles. The family can be further subdivided in a taxonomy. The classification presented here follows Smith (2006),^[1] with the exception of the tribal classification within the Lucaninae.

Subfamily Aesalinae

Tribe Aesalini

Genus Aesalus (Fabricius, 1801)

Aesalus asiaticus Lewis, 1883
Aesalus himalayicus Kurosawa, 1985
Aesalus imanishii Inahara & Ratti, 1991
Aesalus meridionalis Bartolozzi, 1989
Aesalus neotropicalis Bates, 1886
Aesalus saburoi
Aesalus sawaii Fujita & Ichikawa, 1985
Aesalus scarabaeoides Panzer, 1793
Aesalus sichuanensis
Aesalus smithi Bates, 1889
Aesalus trogoides Albers, 1883
Aesalus ulanowskii Ganglbauer, 1886

Genus Cretaesalus (fossil)

Genus Echinoaesalus Zelenka, 1993

Echinoaesalus barriesi (Zelenka, 1993)
Echinoaesalus hidakai (Araya, 1993)
Echinoaesalus jaechi (Zelenka,1993)
Echinoaesalus matsuii (Araya, 1993)
Echinoaesalus sabahensis (Zelenka, 1994)
Echinoaesalus schuhi (Zelenka, 1994)
Echinoaesalus timidus (Krikken, 1975)
Echinoaesalus yongi (Araya, 1993)

Genus Lucanobium Howden & Lawrence, 1974

Lucanobium squamosum Howden & Lawrence, 1974

Tribe Ceratognathini

Genus Ceratognathus Westwood, 1838

Ceratognathus abdominalis Parry, 1870
Ceratognathus bitumulatus Carter, 1925
Ceratognathus flabellatus Boileau, 1905
Ceratognathus frenchi Blackburn, 1897
Ceratognathus froggatti Blackburn, 1894
Ceratognathus gilesi Blackburn, 1895
Ceratognathus macrognathus Boileau, 1905
Ceratognathus mentiferus Westwood, 1863
Ceratognathus minutus Lea, 1929
Ceratognathus niger Westwood, 1838
Ceratognathus ocularis Carter, 1925
Ceratognathus rufipennis Westwood, 1872
Ceratognathus tasmanus Benesh, 1943
Ceratognathus westwoodi Thomson, 1862

Genus Hilophyllus Paulsen & Mondaca, 2006

Hilophyllus argentinensis (Martinez, 1981)
Hilophyllus martinezi Paulsen & Mondaca, 2006
Hilophyllus penai (Martinez, 1976)

Genus Mitophyllus Parry, 1843

Mitophyllus alboguttatus (Bates, 1867)
Mitophyllus dispar (Sharp, 1882)
Mitophyllus foveolatus (Broun, 1880)
Mitophyllus gibbosus (Broun, 1884)
Mitophyllus insignis Broun, 1921
Mitophyllus irroratus Parry, 1842
Mitophyllus macrocerus (Broun, 1886)
Mitophyllus parrianus Westwood, 1863
Mitophyllus reflexus Broun, 1908

Tribe Nicagini

Genus Nicagus LeConte, 1861

Nicagus obscurus (LeConte, 1848)
Nicagus occultus Paulsen & Smith, 2005
Nicagus japonicus Nagel, 1928

Genus Holloceratognathus Nikolajev, 1998

Holloceratognathus cylindricus (Broun, 1895)
Holloceratognathus helotoides (Thomson, 1862)
Holloceratognathus passaliformis (Holloway, 1962)

Subfamily Lampriminae

Genus Dendroblax White, 1846

Dendroblax earlianus White, 1846

Genus Homolamprima

Homolamprima crenulata (MacLeay, 1885)

Genus Lamprima

Lamprima adolphinae (Gestro, 1875)
Lamprima aenea Fabricius, 1792
Lamprima aurata Latreille, 1817
Lamprima imberbis Barber, 1926
Lamprima insularis Macleay, 1885

Genus Phalacrognathus

Phalacrognathus muelleri MacLeay, 1885

Genus Streptocerus Dejean, 1833

Streptocerus speciosus Fairmaire, 1850

Subfamily Lucaninae

Subfamily Lucaninae Latreille, 1804

Tribe Aegini

Genus Aegus MacLeay, 1819

Tribe Allotopini

Genus Allotopus

Allotopus moellenkampi (Fruhstorfer, 1894)
Allotopus rosenbergi (Vollenhoven, 1872)

Genus Mesotopus

Tribe Chiasognathini

Genus Casignetus

Genus Chiasognathus Stephens, 1831

Chiasognathus beneshi Lacroix, 1978
Chiasognathus grantii Stephens, 1831
Chiasognathus jousselini Reiche, 1850
Chiasognathus latreillei Solier, 1851
Chiasognathus mniszechii Kiesche 1919

Genus Protognathinus

Genus Sphaenognathus

Tribe Cladognathini

Genus Aphanognathus

Aphanognathus laterotarsoides (Houlbert, 1915)

Genus Capreolucanus

Capreolucanus sicardi Didier, 1928

Genus Cladophyllus

Genus Eligmodontus

Genus Gonometopus

Genus Macrodorcas Motschulsky, 1862

Macrodorcas adachii
Macrodorcas amamiana
Macrodorcas barbata
Macrodorcas bisignata
Macrodorcas bomansi
Macrodorcas davidi
Macrodorcas dierli
Macrodorcas elsiledis
Macrodorcas fulvonotata
Macrodorcas humilis
Macrodorcas intricata
Macrodorcas kobayashii
Macrodorcas koyamai
Macrodorcas melliana
Macrodorcas mochizukii
Macrodorcas moellenkampi
Macrodorcas montivaga
Macrodorcas pieli
Macrodorcas pseudaxis
Macrodorcas recta Motschulsky, 1858
Macrodorcas ruficrus
Macrodorcas songiana
Macrodorcas striatipennis Motschulsky, 1862
Macrodorcas ursulae
Macrodorcas vernicata
Macrodorcas yaeyamaensis

Genus Palaeognathus

Genus Prismognathus Motschulsky, 1860

Prismognathus alessandrae
Prismognathus angularis Waterhouse, 1874
Prismognathus branczicki
Prismognathus cheni
Prismognathus dauricus Motschulsky, 1860
Prismognathus davidis
Prismognathus delislei
Prismognathus formosanus
Prismognathus klapperichi
Prismognathus lucidus
Prismognathus morimotoi
Prismognathus nigerrimus
Prismognathus nigricolor
Prismognathus piluensis
Prismognathus platycephalus
Prismognathus ruficephalus
Prismognathus sinensis
Prismognathus subnitens
Prismognathus tokui

Genus Prosopocoilus Westwood, 1845

Genus Pseudorhaetus

Genus Rhaetulus

Genus Rhaetus

Rhaetus westwoodi (Parry, 1862)

Genus Tetrarthrius

Genus Weinreichius

Weinreichius perroti Lacroix, 1978

Tribe Colophonini

Genus Colophon

Tribe Cyclommatini

Genus Cyclommatus Parry, 1863

Tribe Dendeziini

Genus Dendezia

Genus Oonotus Parry, 1864

Genus Xiphodontus

Tribe Dorcini

Genus Bartolozziolucanus

Genus Cantharolethrus

Cantharolethrus luxeri

Genus Cyclommatus

Cyclommatus scutellaris

Genus Dorcasoides

Genus Dorcus MacLeay, 1819

Dorcus akahorii
Dorcus alcides
Dorcus alexisi
Dorcus amamianus
Dorcus antaeus
Dorcus anteus
Dorcus arfakianus
Dorcus binodulosus
Dorcus brevis
Dorcus bucephalus
Dorcus carinulatus
Dorcus consentaneus
Dorcus curvidens
Dorcus cylindricus
Dorcus davidis
Dorcus emikoae
Dorcus eurycephalus
Dorcus formosanus
Dorcus gracilicornis
Dorcus garndis
Dorcus hopei
Dorcus hyperion
Dorcus japonicus
Dorcus kyanrauensis
Dorcus metacostatus
Dorcus miwai
Dorcus musimon
Dorcus parallelus
Dorcus parallelipipedus - lesser stag beetle. Linnaeus, 1758
Dorcus parryi
Dorcus parvulus
Dorcus peyronis
Dorcus prochazkai
Dorcus ratiocinativus
Dorcus rectus
Dorcus reichei
Dorcus rudis
Dorcus sawaii
Dorcus schenklingi
Dorcus sewertzowi
Dorcus suturalis
Dorcus taiwanicus
Dorcus tanakai
Dorcus taurus
Dorcus tenuecostatus
Dorcus titanus
Dorcus tormosanus
Dorcus ursulus
Dorcus vavrai
Dorcus velutinus
Dorcus vicinus
Dorcus yamadai

Genus Homoderus

Genus Leptinopterus

Genus Prosopocoilus

Genus Rhaetulus

Rhaetulus crenatus

Genus Serrognathus

Genus Pycnosiphorus

Genus Sclerostomus

Tribe Figulini

Genus Cardanus

Genus Dinonigidius

Genus Figulus

Genus Ganelius

Genus Nigidionus

Genus Nigidius

Genus Novonigidius

Genus Penichrolucanus

Tribe Lissapterini

Genus Bomansius

Genus Dorculus

Genus Geodorcus

Genus Hoplogonus

Genus Lissapterus

Genus Lissotes

Genus Paralissotes

Genus Pseudodorcus

Tribe Lucanini

Genus Hexarthrius Hope, 1842

Hexarthrius aduncusJordan & Rothschild, 1894
Hexarthrius andreasi Schenk, 2003
Hexarthrius bowringii Parry, 1862
Hexarthrius buquettii (Hope, 1843)
Hexarthrius davisoniWaterhouse, 1888
Hexarthrius forsteri (Hope, 1840)
Hexarthrius howdeni De Lisle, 1972
Hexarthrius kirchneri Schenk, 2003
Hexarthrius mandibularis Deyrolle, 1881
Hexarthrius melchioritis Séguy, 1954
Hexarthrius mniszechi(Thomson, 1857)
Hexarthrius nigritus Lacroix, 1990
Hexarthrius parryi Hope, 1842
Hexarthrius rhinoceros(Olivier, 1789)
Hexarthrius vitalisi Didier, 1925

Genus Lucanus

Genus Pseudolucanus

Pseudolucanus barbarossa (Fabricius, 1801)

Tribe Odontolabini

Genus Calcodes

Genus Neolucanus

Neolucanus castanopterus

Genus Odontolabis

Tribe Platycerini

Genus Platyceroides Benesh, 1946

Genus Platyceropsis Benesh, 1946

Platyceropsis keeni (Casey, 1895)

Genus Platycerus Geoffroy, 1762

Tribe Ryssonotini

Genus Cacostomus

Cacostomus floralis (Lea, 1914)
Cacostomus squamosus Newman, 1840
Cacostomus subvittata (Moore, 1994)

Genus Ryssonotus

Subfamily Syndesinae

Tribe Ceruchini

Genus Ceruchus MacLeay, 1819

Ceruchus piceus (Weber, 1801)
Ceruchus punctatus LeConte, 1869
Ceruchus striatus LeConte, 1859
Ceruchus yangi Huang, Imura & Chen, 2011^[2]

Tribe Sinodendronini

Genus Sinodendron Hellwig in Schneider, 1792

Sinodendron cylindricum (Linnaeus, 1758)
Sinodendron rugosum Mannerheim, 1843

Tribe Syndesini

Genus Psilodon Perty, 1830

Psilodon aequinoctiale (Buquet, 1840)
Psilodon gilberti Boucher, 1993
Psilodon schuberti Perty, 1830
Psilodon seguyi (Didier, 1929)
Psilodon xerophilicum Martinez & Reyes-Castillo, 1985

Genus Syndesus MacLeay, 1819

†Syndesus ambericus Woodruff, 2009 (Miocene?; Dominican amber)^[3]
Syndesus cancellatus Montrouzier, 1860
Syndesus cornutus (Fabricius, 1801)
Syndesus macleayi Boileau, 1905

References

↑ Andrew B. T. Smith (2006). "A review of the family-group names for the superfamily Scarabaeoidea (Coleoptera) with corrections to nomenclature and a current classification" (PDF). The Coleopterists Bulletin. 60: 144–204. doi:10.1649/0010-065X(2006)60[144:AROTFN]2.0.CO;2. S2CID 82432509.
↑ Hao Huang, Yuki Imura & Chang-Chin Chen (2011). "Notes on the genus Ceruchus MacLeay from China, with the description of a new species (Coleoptera: Scarabaeoidea: Lucanidae)" (PDF Excerpt). Zootaxa. 2906: 1–42. doi:10.11646/zootaxa.2906.1.1.
↑ Woodruff, R.E. (2009). "A new fossil species of stag beetle from Dominican Republic amber, with Australasian connections (Coleoptera: Lucanidae)". Insecta Mundi. 0098: 1–10.

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[Smith2006-1] Andrew B. T. Smith (2006). "A review of the family-group names for the superfamily Scarabaeoidea (Coleoptera) with corrections to nomenclature and a current classification" (PDF). The Coleopterists Bulletin. 60: 144–204. doi:10.1649/0010-065X(2006)60[144:AROTFN]2.0.CO;2. S2CID 82432509.

[2] Hao Huang, Yuki Imura & Chang-Chin Chen (2011). "Notes on the genus Ceruchus MacLeay from China, with the description of a new species (Coleoptera: Scarabaeoidea: Lucanidae)" (PDF Excerpt). Zootaxa. 2906: 1–42. doi:10.11646/zootaxa.2906.1.1.

[Woodruff2009-3] Woodruff, R.E. (2009). "A new fossil species of stag beetle from Dominican Republic amber, with Australasian connections (Coleoptera: Lucanidae)". Insecta Mundi. 0098: 1–10.

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