Tylototriton
Tylototriton shanjing
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Amphibia
Order: Urodela
Family: Salamandridae
Subfamily: Pleurodelinae
Genus: Tylototriton
Andersson, 1871
Type species
Tylototriton verrucosus
Anderson, 1871

Tylototriton is a genus of newts known as crocodile newts or knobby newts. About 36 known species are in this genus.[1][2][3] Many species have been described just recently. They range from northeastern India and Nepal through Burma to northern Thailand, Laos, Vietnam, and southern China.[2]

Species

38 species recognized as of July 2023:[4]

  • Tylototriton anguliceps Le, Nguyen, Nishikawa, Nguyen, Pham, Matsui, Bernardes, and Nguyen, 2015
  • Tylototriton anhuiensis Qian, Sun, Li, Guo, Pan, Kang, Wang, Jiang, Wu, and Zhang, 2017
  • Tylototriton asperrimus Unterstein, 1930
  • Tylototriton broadoridgus Shen, Jiang, and Mo, 2012
  • Tylototriton dabienicus Chen, Wang, and Tao, 2010
  • Tylototriton daloushanensis Zhou, Xiao, and Luo, 2022
  • Tylototriton hainanensis Fei, Ye, and Yang, 1984
  • Tylototriton himalayanus Khatiwada, Wang, Ghimire, Vasudevan, Paudel, and Jiang, 2015
  • Tylototriton kachinorum Zaw, Lay, Pawangkhanant, Gorin, and Poyarkov, 2019
  • Tylototriton kweichowensis Fang and Chang, 1932
  • Tylototriton liuyangensis Yang, Jiang, Shen, and Fei, 2014
  • Tylototriton lizhengchangi Hou, Zhang, Jiang, Li and Lu, 2012
  • Tylototriton maolanensis Li, Wei, Cheng, Zhang, and Wang, 2020
  • Tylototriton ngarsuensis Grismer, Wood, Quah, Thura, Espinoza, Grismer, Murdoch, and Lin, 2018
  • Tylototriton ngoclinhensis Phung, Pham, Nguyen, Ninh, Nguyen, Bernardes, Le, Ziegler & Nguyen, 2023
  • Tylototriton notialis Stuart, Phimmachak, Sivongxay, and Robichaud, 2010
  • Tylototriton panhai Nishikawa, Khonsue, Pomchote, and Matsui, 2013
  • Tylototriton panwaensis Grismer, Wood, Quah, Thura, Espinoza, and Murdoch, 2019
  • Tylototriton pasmansi Bernardes, Le, Nguyen, Pham, Pham, Nguyen, and Ziegler, 2020
  • Tylototriton phukhaensis Pomchote, Khonsue, Thammachoti, Hernandez, Peerachidacho, Suwannapoom, Onishi, and Nishikawa, 2020
  • Tylototriton podichthys Phimmachak, Aowphol, and Stuart, 2015
  • Tylototriton pseudoverrucosus Hou, Gu, Zhang, Zeng, and Lu, 2012
  • Tylototriton pulcherrimus Hou, Zhang, Li, and Lu, 2012
  • Tylototriton shanjing Nussbaum, Brodie, and Yang, 1995
  • Tylototriton shanorum Nishikawa, Matsui, and Rao, 2014
  • Tylototriton sini Lyu, Wang, Zeng, Zhou, Qi, Wan, and Wang, 2021
  • Tylototriton sparreboomi Bernardes, Le, Nguyen, Pham, Pham, Nguyen, and Ziegler, 2020
  • Tylototriton taliangensis Liu, 1950
  • Tylototriton thaiorum Poyarkov, Nguyen, and Arkhipov, 2021
  • Tylototriton tongziensis Li, Liu, Shi, Wei, and Wang, 2022
  • Tylototriton umphangensis Pomchote, Peerachidacho, Hernandez, Sapewisut, Khonsue, Thammachoti, and Nishikawa, 2021
  • Tylototriton uyenoi Nishikawa, Khonsue, Pomchote, and Matsui, 2013
  • Tylototriton verrucosus Anderson, 1871
  • Tylototriton vietnamensis Böhme, Schöttler, Nguyen, and Köhler, 2005
  • Tylototriton wenxianensis Fei, Ye, and Yang, 1984
  • Tylototriton yangi Hou, Zhang, Zhou, Li, and Lu, 2012
  • Tylototriton zaimeng Decemson, Lalremsanga, Elangbam, Vabeiryureilai, Shinde, Purkayastha, Arkhipov, Bragin & Poyarkov, 2023
  • Tylototriton ziegleri Nishikawa, Matsui, and Nguyen, 2013

Phylogeny

The following phylogeny of Tylototriton is from Nishikawa, Matsui & Rao (2014).[5]

Tylototriton 
Subgenus Yaotriton 

Tylototriton panhai

Tylototriton asperrimus

Subgenus Tylototriton 

Tylototriton taliangensis

Tylototriton kweichowensis

Tylototriton verrucosus verrucosus (Nepal)

Tylototriton verrucosus verrucosus (Myanmar)

Tylototriton yangi

Tylototriton uyenoi

T. shanjing complex 

Tylototriton verrucosus pulcherrima

Tylototriton shanjing

Tylototriton verrucosus verrucosus

References

  1. ↑ Max Sparreboom (2013). "Tylototriton Anderson, 1871". Salamanders of the Old World. Retrieved 14 April 2013.
  2. 1 2 Frost, Darrel R. (2021). "Tylototriton Anderson, 1871". Amphibian Species of the World 6.1, an Online Reference. American Museum of Natural History. Retrieved 19 November 2021.
  3. ↑ Nishikawa, Kanto; Masafumi Matsui; Tao Thien Nguyen (2013). "A new species of Tylototriton from northern Vietnam (Amphibia: Urodela: Salamandridae)". Current Herpetology. 32: 34–49. doi:10.5358/hsj.32.34. S2CID 59036508.
  4. ↑ Phung, Trung My; Pham, Cuong The; Nguyen, Truong Quang; Ninh, Hoa Thi; Nguyen, Huy Quoc; Bernardes, Marta; Le, Son Thanh; Ziegler, Thomas; Nguyen, Tao Thien (2023-07-03). "Southbound – the southernmost record of Tylototriton (Amphibia, Caudata, Salamandridae) from the Central Highlands of Vietnam represents a new species". ZooKeys (1168): 193–218. doi:10.3897/zookeys.1168.96091. ISSN 1313-2970.
  5. ↑ Nishikawa, Matsui & Rao, 2014 : A new species of Tylototriton (Amphibia: Urodela: Salamandridae) from central Myanmar. Natural History Bulletin of the Siam Society, vol. 60, no. 1, p. 9–22.
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