Tyrannotitan Temporal range: Early Cretaceous (Aptian),
Reconstructed skeleton in Queensland Museum
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Clade:	Dinosauria
Clade:	Saurischia
Clade:	Theropoda
Family:	†Carcharodontosauridae
Tribe:	†Giganotosaurini
Genus:	†Tyrannotitan Novas et al., 2005
Type species
†Tyrannotitan chubutensis Novas et al., 2005

Tyrannotitan (/tɪˌrænəˈtaɪtən/; lit. 'tyrant titan') is a genus of large bipedal carnivorous dinosaur of the carcharodontosaurid family from the Aptian stage of the early Cretaceous period, discovered in Argentina. It is closely related to other giant predators like Carcharodontosaurus and especially Giganotosaurus as well as Mapusaurus.

Discovery and species

Known remains (in yellow)

Vertebra and ischium

Tyrannotitan chubutensis was described by Fernando E. Novas, Silvina de Valais, Pat Vickers-Rich, and Tom Rich in 2005. The fossils were found at La Juanita Farm, 28 kilometres (17 mi) northeast of Paso de Indios, Chubut Province, Argentina. They are believed to have been from the Cerro Castaño Member, Cerro Barcino Formation (Aptian stage).^[1]

The holotype material was designated MPEF-PV 1156 and included partial dentaries, teeth, back vertebrae 3–8 and 11–14, proximal tail vertebrae, ribs and chevrons, a fragmentary scapulocoracoid, humerus, ulna, partial ilium, a nearly complete femur, fibula, and left metatarsal 2. Additional material (designated MPEF-PV 1157) included jugals, a right dentary, teeth, atlas vertebra, neck vertebra (?) 9, back vertebrae (?)7, 10, 13, fused sacral centra (5 total), an assortment of distal caudals, ribs, the right femur, a fragmentary left metatarsal 2, pedal phalanges 2-1, 2–2, and 3-3.^[1]

Description

Reconstruction of a Tyrannotitan, feeding on a Chubutisaurus.

Estimated size, compared to a human.

Tyrannotitan was a large animal, reaching 12.2–13 metres (40–43 ft) in length and 4.8–7 t (5.3–7.7 short tons) in body mass.^{[2][3][4][5][6]} Unlike other known carcharodontosaurids, this animal lacks skeletal pneumaticity extending into the sacral and caudal centra. The scapulocoracoid is fused, and much better developed than that of Giganotosaurus carolinii, yet the arm is very small. Most of the shaft of the scapula is missing.^[1]

The acromion curves about 90 degrees from the shaft axis, making it look vaguely tyrannosaurid-like. Whether the sharp difference between taxa is due to evolution or sexual dimorphism in poorly sampled populations of both species, has not been determined (the latter seems unlikely). A proximal caudal has a very tall neural spine (about twice the height of its centrum, judging by the figure). The base of the orbital fenestra is a notch of nearly 90 degrees into the body of the jugal, which contrasts with the rounded base restored for Giganotosaurus and agrees with Carcharodontosaurus favorably. The denticles on its teeth are "chisel-like", and are virtually identical to those of other carcharodontosaurids in having a wrinkled enamel surface, heavily serrated mesial and distal carinae, and labiolingually compressed (laterally flattened) crowns.^[7] The femur of the paratype specimen is 1.4 m (4.6 ft) long according to Novas et al.^[1] Canale et al. recover Tyrannotitan as deeply nested within the tribe Giganotosaurini as its most basal member. Characteristics that unite the Giganotosaurini include the presence of a postorbital process on the jugal with a wide base, and a derived femur with a weak fourth trochanter and a shallow broad extensor groove at the distal end.^[7][8]

Classification

The following cladogram after Novas et al., 2013, shows the position of Tyrannotitan within Carcharodontosauridae.^[9]

References